George Zug

• B.S., Albright College, 1960
• M.S., University of Florida, 1963
• Ph.D., University of Michigan, 1968

Herpetology/biology and systematics of amphibians and reptiles; currently focused on the systematics of the herpetofauna of Oceania and Burma.

Book

• Reese, Devin A. and Zug, George R. 2023. Bibliography of Tortoises of the World Giants to Dwarfs. Washington, DC: Division of Amphibians and Reptiles, National Museum of Natural History. In Smithsonian Herpetological Information Service, 161.
• Zug, George R. 2022. Amphibians and Reptiles of Myanmar: Checklist and Keys. I. Amphibians, Crocodilians, and Turtles. Washington, DC: Smithsonian Scholarly Press. In Smithsonian Contributions to Zoology, 653. https://doi.org/10.5479/si.19098995.
• Zug, George R. and Mulcahy, Daniel G. 2020. Identification Guide Amphibians & Reptiles of South Tanintharyi. Yangon: Fauna & Flora International.
• Zug, George R. 2013. Reptiles and Amphibians of the Pacific Islands: a comprehensive guide. Berkeley: University of California Press.
• Zug, George R. 2013. Literature resources For “Reptiles And Amphibians Of The Pacific Islands: a comprehensive guide”. In Smithsonian Herpetological Information Service, 143. https://doi.org/10.5479/si.23317515.143.1.
• Zug, George R. 2012. Data resources for Tropical Asian Dry Forest Amphibians and Reptiles. Smithsonian Institution. In Smithsonian Herpetological Information Service, 141.
• Zug, George R. 2010. Speciation and dispersal in a low diversity taxon: the Slender Geckos Hemiphyllodactylus (Reptilia, Gekkonidae). In Smithsonian Contributions to Zoology, 631. https://doi.org/10.5479/si.00810282.631.
• Leviton, A. E., Zug, George R., Vindum, J. V., and Wogan, Guinevere O. U. 2008. Handbook to the Dangerously Venomous Snakes of Myanmar. San Francisco: California Academy of Sciences.
• Zug, George R. and Ernst, Carl H. 2004. Snakes: Smithsonian Answer Book. Smithsonian Books.
• Slowinski, Joseph B. and Zug, George R. 2000. Presentation - Herpetological surveys in Myanmar.
• Zug, George R. 1999. Preparation of voucher specimins: amphibians and reptiles.
• Zug, George R. 1999. Preparation of voucher specimens: Amphibians and reptiles.
• Mitchell, Joseph C. and Zug, George R. 1995. Keys to the known amphibians and reptiles of the Royal Chitwan National Park, Nepal. In Smithsonian Herpetological Information Service, 106. https://doi.org/10.5479/si.23317515.106.1.
• Zug, George R., Wynn, Addison H., and Ruckdeschel, Carol. 1986. Age determination of loggerhead sea turtles, Caretta caretta, by incremental growth marks in the skeleton. Washington, D.C.: Smithsonian Institution Press. In Smithsonian Contributions to Zoology, 427. https://doi.org/10.5479/si.00810282.427.
• Irish, F. J. and Zug, George R. 1982. Biography and bibliography of James A. Peters. In Smithsonian Herpetological Information Service, 51. https://doi.org/10.5479/si.23317515.51.1.
• Zug, George R., Hedges, S. Blair, and Sunkel, Sara. 1979. Variation in reproductive parameters of three neotropical snakes, Coniophanes fissidens, Dipsas catesbyi, and Imantodes cenchoa. Washington: Smithsonian Institution Press. In Smithsonian Contributions to Zoology, 300. https://doi.org/10.5479/si.00810282.300.
• Zug, George R. and Zug, Patricia B. 1979. The marine toad, Bufo marinus : a natural history resumé of native populations. Washington: Smithsonian Institution Press. In Smithsonian Contributions to Zoology, 284. https://doi.org/10.5479/si.00810282.284.
• Zug, George R. 1978. Anuran locomotion-structure and function. Washington: Smithsonian Institution Press. In Smithsonian Contributions to Zoology, 276. https://doi.org/10.5479/si.00810282.276.
• Zug, George R. 1971. The distribution and patterns of the major arteries of the Iguanids and comments on the intergeneric relationships of Iguanids (Reptilia: Lacertilia). Washington: Smithsonian Institution Press; for sale by the Supt. of Docs., U.S. Govt. Print. Off.. In Smithsonian Contributions to Zoology, 83. https://doi.org/10.5479/si.00810282.83.
• Jacobs, George, Zug, George R., and Peters, James Arthur. 1970. Herpetology - by I.S. Darevskii. In Smithsonian Herpetological Information Service, 22.
• Zug, George R. 1970. Index. The Salamanders Of The Family Plethodontidae (by Emmett Reid Dunn). In Smithsonian Herpetological Information Service, 19. https://doi.org/10.5479/si.23317515.19.

Journal Article

• Mulcahy, Daniel G., Ibáñez, Roberto D., Jaramillo, Cesar A., Crawford, Andrew J., Ray, Julie M., Gotte, Steve W., Jacobs, Jeremy F., Wynn, Addison H., Gonzalez-Porter, Gracia, McDiarmid, Roy W., Crombie, Ronald I., Zug, George R., and de Queiroz, Kevin. 2022. "DNA barcoding of the National Museum of Natural History reptile tissue holdings raises concerns about the use of natural history collections and the responsibilities of scientists in the molecular age." PLOS ONE, 17, (3). https://doi.org/10.1371/journal.pone.0264930.
• Miller, Aryeh H., Zug, George R., Wogan, Guinevere O. U., Lee, Justin L., and Mulcahy, Daniel G. 2021. "Phylogeny, Diversity, and Distribution of Micryletta (Anura: Microhylidae) in Myanmar." Ichthyology & Herpetology, 109, (1) 245–257. https://doi.org/10.1643/h2020100.
• Avens, Larisa, Goshe, Lisa R., Zug, George R., Balazs, George H., Benson, Scott R., and Harris, Heather. 2019. "Regional comparison of leatherback sea turtle maturation attributes and reproductive longevity." Marine Biology, 167, (1) Article 4. https://doi.org/10.1007/s00227-019-3617-y.
• Lee, Justin L., Miller, Aryeh H., Zug, George R., and Mulcahy, Daniel G. 2019. "The discovery of Rock Geckos Cnemaspis Strauch, 1887 (Squamata: Gekkonidae) in the Tanintharyi Region, Myanmar with the description of two new species." Zootaxa, 4661, (1) 40–64. https://doi.org/10.11646/zootaxa.4661.1.2.
• Grismer, L. L., Wood, Perry L.,Jr, Zug, George R., Thura, Myint Kyaw, Grismer, Marta S., Murdoch, Matthew L., Quah, Evan S. H., and Lin, Aung. 2018. "Two more new species of Hemiphyllodactylus Bleeker (Squamata: Gekkonidae) from the Shan Hills of eastern Myanmar (Burma)." Zootaxa, 4483, (2) 295–316. https://doi.org/10.11646/zootaxa.4483.2.4.
• Lee, Justin L., Miller, Aryeh H., Connette, Grant M., Oo, Khin Swe, Zug, George R., and Mulcahy, Daniel G. 2018. "First record of the Malaysian Bridle Snake, Dryocalamus subannulatus (Duméril, Bibron & Duméril, 1854), in Myanmar (Reptilia, Serpentes, Colubridae)." Check List, 14, (2) 341–345. https://doi.org/10.15560/14.2.341.
• Lee, Justin L., Vogel, Gernot, Miller, Aryeh H., and Zug, George R. 2018. "Rediscovery of Xenochrophis bellulus (Stoliczka, 1871) from Myanmar (Serpentes: Natricinae) with comments on its taxonomic status." Proceedings of the Biological Society of Washington, 131, (1) 19–35. https://doi.org/10.2988/17-00025.
• Mulcahy, Daniel G., Lee, Justin L., Miller, Aryeh H., Chand, Mia, Thura, Myint Kyaw, and Zug, George R. 2018. "Filling the BINs of life: Report of an amphibian and reptile survey of the Tanintharyi (Tenasserim) Region of Myanmar, with DNA barcode data." ZooKeys, 757 85–152. https://doi.org/10.3897/zookeys.757.24453.
• Zug, George R., Mulcahy, Daniel G., Vindum, Jens V., and Wilkinson, Jeffery A. 2018. "Anurans of the Tanintharyi Region, Myanmar: a preview of increasing species diversity." Alytes, 36, (1–4) 266–275.
• Connette, Grant M., Oswald, Patrick, Thura, Myint Kyaw, LaJeunesse Connette, Katherine J., Grindley, Mark E., Songer, Melissa A., Zug, George R., and Mulcahy, Daniel G. 2017. "Rapid forest clearing in a Myanmar proposed national park threatens two newly discovered species of geckos (Gekkonidae: Cyrtodactylus)." PLOS ONE, 12, (4) 1–18. https://doi.org/10.1371/journal.pone.0174432.
• Mulcahy, Daniel G., Lee, Justin L., Miller, Aryeh H., and Zug, George R. 2017. "Troublesome Trimes: Potential cryptic speciation of the Trimeresurus (Popeia) popeiorum complex (Serpentes: Crotalidae) around the Isthmus of Kra (Myanmar and Thailand)." Zootaxa, 4347, (2) 301–315. https://doi.org/10.11646/zootaxa.4347.2.6.
• Zug, George R., Mulcahy, Daniel G., and Vindum, Jens V. 2017. "Resurrection of Bronchocela burmana Blanford, 1878 for the Green Crested Lizard (Squamata, Agamidae) of southern Myanmar." ZooKeys, 657 141–156. https://doi.org/10.3897/zookeys.657.11600.
• Miller, Aryeh H. and Zug, George R. 2016. "Morphology and Biology of the Asian Common Mockviper, Psammodynastes pulverulentus (Serpentes: Lamprophiidae): a focus on Burmese populations." Proceedings of the Biological Society of Washington, 129, (1) 173–194. https://doi.org/10.2988/0006-324X-129.Q2.173.
• Miller, Aryeh H. and Zug, George R. 2016. "An enigmatic ^Lygosoma^ (Reptilia: Squamata: Scincidae) from northern Myanmar." Herpetological Review, 47, (3) 373–374.
• Lee, Justin L., Myint, Kyaw Thura, Mulcahy, Daniel G., and Zug, George R. 2015. "Three Colubrid Snakes New To Myanmar." Herpetology Notes, 8 217–220.
• Zug, George R. 2015. "Morphology and systematics of Kalophrynus interlineatus-pleurostigma populations (Anura, Microhylidae, Kalophryninae) and a taxonomy of the genus Kalophrynus Tschudi, Asian Sticky Frogs." Proceedings of the California Academy of Sciences, 62, (5) 135–190.
• Mulcahy, Daniel G., Martínez-Gómez, Juan E., Aguirre-León, Gustavo, Cervantes-Pasqualli, Juan, and Zug, George R. 2014. "Rediscovery of an Endemic Vertebrate from the Remote Islas Revillagigedo in the Eastern Pacific Ocean: The Clarión Nightsnake Lost and Found." PLoS ONE, 9, (5) 1–8. https://doi.org/10.1371/journal.pone.0097682.
• Nguyen, Truong Quang, Botov, Andreas, Le, Minh Duc, Nophaseud, Liphone, Zug, George R., Bonkowski, Michael, and Ziegler, Thomas. 2014. "A new species of Hemiphyllodactylus (Reptilia: Gekkonidae) from northern Laos." Zootaxa, 3827, (1) 45–56.
• Zug, George R. 2014. "Comment on the proposed confirmation of the availability of Spracklandus Hoser, 2009 (Reptilia, Squamata, Elapidae)." The Bulletin of Zoological Nomenclature, 71, (4) 252–253.
• Zug, George R. and Kaiser, Hinrich. 2014. "A new species of four-toed skink (Squamata: Scincidae: Carlia peronii species group) from Pulau Sukur, Indonesia, and biogeographic notes on the herpetofauna of Flores and Komodo." Proceedings of the Biological Society of Washington, 126, (4) 379–392. https://doi.org/10.2988/0006-324X-126.4.379.
• Böhm, Monika, Collen, Ben, Baillie, Jonathan E. M., Bowles, Philip, Chanson, Janice, Cox, Neil, Hammerson, Geoffrey, Hoffmann, Michael, Livingstone, Suzanne R., Ram, Mala, Rhodin, Anders G. J., Stuart, Simon N., van Dijk, Peter Paul, Young, Bruce E., Afuang, Leticia E., Aghasyan, Aram, García, Andrés, Aguilar, César, Ajtic, Rastko, Akarsu, Ferdi, Alencar, Laura R. V., Allison, Allen, Ananjeva, Natalia, Anderson, Steve, Andrén, Claes et al. 2013. "The conservation status of the world's reptiles." Biological Conservation, 157 372–385. https://doi.org/10.1016/j.biocon.2012.07.015.
• Fisher, Robert N., Uili, Moeumu, Enoka, Fialelei, and Zug, George R. 2013. "Clarifying the distributional records for Hemiphyllodactylus typus (Bleeker, 1860) from Central Oceania." Herpetology Notes, 6 247–250.
• Zug, George R. and Kaiser, Hinrich. 2013. "Kalophrynus Tschudi, 1838 (Amphibia, Anura, Microhylidae): proposed conservation by designation of a neotype for its type species Kalophrynus pleurostigma Tschudi, 1838." The Bulletin of Zoological Nomenclature, 70, (2) 86–88.
• Zug, George R. and Kaiser, Hinrich. 2013. "Comment on the proposed conservation of Kalophrynus Tschudi, 1838 (Amphibia, Anura, Microhylidae) by designation of a neotype for its type species Kalophrynus pleurostigma Tschudi, 1838." The Bulletin of Zoological Nomenclature, 71, (1) 28.
• Zug, George R. 2012. "A new species of treeskink (Squamata: Scincidae: Emoia samoensis species group) from Rotuma, south-central Pacific." Proceedings of the Biological Society of Washington, 125, (1) 74–84. https://doi.org/10.2988/11-24.1.
• Zug, George R. 2012. "Lizards of Tonga with a description of a new Tongan treeskink (Squamata: Scincidae: Emoia samoensis Group)." Pacific Science, 66, (2) 225–237. https://doi.org/10.2984/66.2.9.
• Zug, George R. and Fisher, R. N. 2012. "A preliminary assessment of the Nactus pelagicus species group (Squamata: Gekkonidae) in New Guinea and a new species from the Admiralty Islands." Zootaxa, 3257 22–37.
• Austin, Christopher C., Rittmeyer, Eric N., Oliver, Lauren A., Andermann, John O., Zug, George R., Rodda, Gordon H., and Jackson, Nathan D. 2011. "The bioinvasion of Guam: inferring geographic origin, pace, pattern and process of an invasive lizard (Carlia) in the Pacific using multi-locus genomic data." Biological Invasions, 13, (9) 1951–1967. https://doi.org/10.1007/s10530-011-0014-y.
• Zug, George R., Gotte, Steve W., and Jacobs, Jeremy F. 2011. "Pythons in Burma: Short-tailed Pythons (Reptilia: Squamata)." Proceedings of the Biological Society of Washington, 124, (2) 112–136. https://doi.org/10.2988/10-34.1.
• Zug, George R., Hamilton, Alison M., and Austin, Christopher C. 2011. "A new Emoia samoensis group lizard (Squamata: Scincidae) from the Cook Islands, South-central Pacific." Zootaxa, 2765 47–57.
• Austin, Christopher C., Rittmeyer, Eric N., Richards, Stephen J., and Zug, George R. 2010. "Molecular phylogeny, historical biogeography and body size evolution in Pacific crocodile skinks Tribolonotus (Squamata; Scincidae)." Molecular Phylogenetics and Evolution, 57 227–236.
• Balmer, Oliver, Ciofi, Claudio, Galbraith, David A., Swingland, Ian R., Zug, George R., and Caccone, Adalgisa. 2010. "Population genetic structure of Aldabra giant tortoises." The Journal of Heredity, 102, (1) 29–37. https://doi.org/10.1093/jhered/esq096.
• Furness, Andrew I., McDiarmid, Roy W., Heyer, W. Ronald, and Zug, George R. 2010. "Oviduct modifications in foam-nesting frogs, with emphasis on the genus Leptodactylus (Amphibia, Leptodactylidae)." South American Journal of Herpetology, 5, (1) 13–29. https://doi.org/10.2994/057.005.0102.
• Hamilton, Alison M., Zug, George R., and Austin, Christopher C. 2010. "Biogeographic anomaly or human introduction: a tree skink (Reptilia: Squamata) in the Cook Islands?" Biological Journal of the Linnean Society, 100 318–328.
• Watling, Dick, Wynn, Addison H., and Zug, George R. 2010. "Rediscovery of the Taveuni blind snake." Oryx, 44, (2) 165–166.
• Zug, George R. 2010. "An Outlying Carlia Population from Java and Comments on Species Groups within the Genus Carlia (Reptilia: Squamata: Scincidae)." Proceedings of the California Academy of Sciences, 61, (8) 389–408.
• Zug, George R., Jacobs, Jeremy F., Vindum, Jens V., and Win, Kyi Soe. 2010. "Non-reproductive seasonal colour change in a population of Calotes "versicolor" from Myanmar (Squamata: Agamidae)." Salamandra, 46, (2) 104–107.
• Curtin, A. J., Zug, George R., and Spotila, J. R. 2009. "Longevity and growth strategies of the desert tortoise (Gopherus agassizii) in two American deserts." Journal of Arid Environments, 73, (4-5) 463–471. https://doi.org/10.1016/j.jaridenv.2008.11.011.
• Zug, George R. 2009. "Comments on the proposed conservation of usage of Testudo gigantea Schweigger, 1812 (currently Geochelone (Aldabrachelys) gigantea; Reptilia, Testudines) (Case 3463)." The Bulletin of Zoological Nomenclature, 66, (1) 80–80.
• Zug, George R., Vindum, Jens J., Jacobs, Jeremy F., Lwin, Kyi Soe, and Win, Ye Myo. 2009. "Calotes "versicolor" (Burmese garden lizard). Gliding." Herpetological Review, 40, (3) 340–341.
• Curtin, A. J., Zug, George R., Medic, P. A., and Spotila, J. R. 2008. "Assessing age in the desert tortoise Gopherus agassizii: testing skeletochronology with individuals of known age." Endangered Species Research, 5, (1) 21–27. https://doi.org/10.3354/esr00108.
• Mahony, Stephen and Zug, George R. 2008. "Hemidactylus karenorum (Squamata, Gekkonidae) in India." Hamadryad, 32, (1) 73–75.
• McMahan, Caleb D. and Zug, George R. 2007. "Burmese Hemidactylus (Reptilia, Squamata, Gekkonidae): geographic variation in the morphology of Hemidactylus bowringii in Myanmar and Yunnan." Proceedings of the California Academy of Sciences, 58, (24) 485–509.
• Zug, George R., Balazs, George H., Wetherall, Jerry A., Parker, Denise M., and Murakawa, Shawn K. K. 2007. "Age and growth of Hawaiian green seaturtles (Chelonia mydas): an analysis based on skeletochronology." Fishery Bulletin, 117–127.
• Zug, George R., Vindum, Jens J., and Koo, Michelle. 2007. "Burmese Hemidactylus (Reptilia, Squamata, Gekkonidae): taxonomic notes on tropical Asian Hemidactylus." Proceedings of the California Academy of Sciences, 58, (19) 387–405.
• Zug, George R. 2006. "Lepidodactylus (Squamata: Gekkonidae) in Islands Asia: A L. aureolineatus from Sulawesi." Hamadryad, 30, (1) 211–213.
• Zug, George R. and Allison, Allen. 2006. "New Carlia fusca complex lizards (Reptilia : Squamata : Scincidae) from New Guinea, Papua-Indonesia." Zootaxa, (1237) 27–44.
• Zug, George R., Blackburn, John L., and Kyi, Sai Wunna. 2006. "Checkered Keelbacks (Xenochrophis-Reptilia: Serpentes: Natricidae) at the Moyingyi Wetland Bird Sanctuary, Myanmar." Hamadryad, 30, (1) 157–166.
• Zug, George R., Brown, K. H., Schulte, James A., II, and Vindum, J. V. 2006. "Systematics of the gardern lizards, Calotes versicolor group (Reptilia, Squamata, Agamidae), in Myanmar: Central Dry Zone populations." Proceedings of the California Academy of Sciences, 57, (2) 35–68.
• Zug, George R., Chaloupka, Milani, and Balazs, George H. 2006. "Age and Growth in Olive Ridley Seaturtles (Lepidochelys olivacea) from the North-Central Pacific: A Skeletochronological Analysis." Marine Ecology, 27 263–270.
• Couper, Patrick J., Wilmer, Jessica Worthington, Roberts, Lewis, Amey, Andrew P., and Zug, George R. 2005. "Skinks Currently Assigned to Carlia aerata (Scincidae: Lygosominae) of North-Eastern Queensland: A Preliminary Study of Cryptic Diversity and Two New Species." Australian Journal of Zoology, 53, (1) 35–49.
• Zug, George R., Kyi, Sai Wunna, and Win, Htun. 2005. "Turtles in Shwesettaw Wildlife Sanctuary, Myanmar." Turtle and Tortoise Newsletter, (8) 4–4.
• Zug, George R. 2004. "Amphibians and Reptiles of Nepal: Biology, Systematics, Field Guide, ed. Schliech & KAstle." Herpetological Review, 35, (1) 88–90.
• Zug, George R. 2004. "Systematics of the Carlia "fusca" Lizards (Squamata: Scincidae) of New Guinea and Nearby Islands." Bishop Museum Bulletin, 5 1–84.
• Glesenkamp, L., Zug, George R., and Mitchell, J. 2003. "Reproductive cycle of male snapping turtles (Chelydra serpentina) in southeastern Virginia." Chelonian Conservation and Biology, 4, (3) 697–700.
• Kyi, Sai Wunna and Zug, George R. 2003. "Unusual Foraging Behavior of Naja kaouthia at the Moyingue Wetlands Birds Sanctuary, Myanmar." Hamadryad, 27, (2) 265–266.
• Leviton, Alan E., Wogan, G., Koo, M., Zug, George R., Lucas, R., and Vindum, J. 2003. "The dangerously venomous snakes of Myanmar. Illustrated key and checklist." Proceedings of the California Academy of Sciences, 54 407–462.
• Zug, George R., Watling, D., Alefaio, T., Alefaio, S., and Ludescher, C. 2003. "A new gecko (Lepidodactylus: Reptilia: Squamata) from Tuvalu, south-central Pacific." Proceedings of the Biological Society of Washington, 116, (1) 38–46.
• McDiarmid, Roy W., de Queiroz, Kevin, Beaman, Kent, Crother, Brian, Etheridge, Richard, Flores Villela, Oscar, Frost, Darrel, Grismer, L. Lee, Hollingsworth, Bradford D., Kearney, Maureen, McGuire, Jimmy A., Wright, John, and Zug, George R. 2002. "Comment on the proposed precedence of the specific name of Euphryne obesus Baird, 1859 over that of Sauromalus ater Dumeril, 1856 (Reptilia, Squamata)." The Bulletin of Zoological Nomenclature, 59, (1) 45–48.
• Schmalz, Timothy D. and Zug, George R. 2002. "Observations on Geographic Variation in the Asian Frog, Hoplobatrachus rugulosus (Anura: Ranidae)." Hamadryad, 27, (1) 90–98.
• Zug, George R. and Balazs, George H. 2000. "Estimating Age in Hawaiian Green Sea Turtles by Skeletochronology." NOAA Technical Memorandum NMFS-SEFSC, 436 127–128.
• Austin, C. C. and Zug, George R. 1999. "Molecular and Morphological Evolution in the South-Central Pacific Skink Emoia tongana (Reptilia: Squamata): Uniformity and Human-mediated Dispersal." Australian Journal of Zoology, 47 425–437.
• Ota, H., Fisher, R., Ineich, I., Case, T., Radtkey, R., and Zug, George R. 1998. "A New Lepidodactylus (Squamata: Gekkonidae) from Vanuatu." Herpetologica, 54, (3) 325–332.
• Zug, George R. 1998. "Galápagos Tortoise Nomenclature: Still Unresolved." Chelonian Conservation Biology, 2, (4) 618–619.
• Zug, George R. 1998. "Australian Populations of the Nactus pelagicus Complex (Reptilia: Gekkonidae)." Memoirs Queensland Museum, 42, (2) 613–626.
• Zug, George R. and Glor, R. E. 1998. "Estimates of age and growth in a population of green sea turtles (Chelonia mydas) from the Indian River lagoon system, Florida: a skeletochronological analysis." Canadian Journal of Zoology, 76, (8) 1497–1506.
• Zug, George R., Win, H., Thin, T., Min, T. Z., Lhon, W. Z., and Kyaw, K. 1998. "Herpetofauna of the Chatthin Wildlife Sanctuary, North-Central Myanmar with preliminary observations of their natural history." Hamadryad, 23, (2) 111–120.
• Chaloupka, M. and Zug, George R. 1997. "A polyphasic growth function for the endangered Kemp's ridley sea turtle, Lepidochelys kempii." Fishery Bulletin, 95, (4) 849–856.
• Parham, James F. and Zug, George R. 1997. "Age and Growth of Loggerhead Sea Turtles (Caretta Caretta) of Coastal Georgia: An Assessment of Skeletochronogical Age-Estimates." Bulletin of Marine Science, 61, (2) 287–304.
• Zug, George R. 1997. "Striped Skinks of Oceania: the Status of Emoia caeruleocauda in Fiji." Pacific Science, 51, (2) 183–188.
• Zug, George R. 1997. "Review. Tadpoles." Science Books & Films, 33, (2) 52.
• Zug, George R. 1997. "Look Inside Reptiles (review)." Science Activities, 33, (3) 44.
• Zug, George R. and Gill, B. J. 1997. "Morphological variation of Emoia murphyi (Lacertilia: Scincidae) on islands of the southwest Pacific." Journal of the Royal Society of New Zealand, 27, (2) 235–242.
• Zug, George R. and Ineich, I. 1997. "The Tongan Giant Lizard Tachygyia. Extinct or Extant?" Cryptozoology, 12 30–35.
• Zug, George R., Kalb, Heather J., and Luzar, Stephen J. 1997. "Age and Growth in Wild Kemp's Ridley Seaturtles Lepidochelys kempii from Skeletochronological Data." Biological Conservation, 80 261–268.
• Ineich, Ivan and Zug, George R. 1996. "Tachygyia, the giant Tongan skink: extinct or extant?" Cryptozoology, 12 30–35.
• Parham, J. F., Dodd, C. K., and Zug, George R. 1996. "Skeletochronological age estimates for the Red Hills salamander, Phaeognathus hubrichti." Journal of Herpetology, 30, (3) 401–404.
• Parham, James F. and Zug, George R. 1996. "Chelonia agassizii —Valid or Not?" Marine Turtle Newsletter, (72) 2–4.
• Zug, George R. and Parham, J. F. 1996. "Age and Growth in Leatherback Turtles, Dermochelys coriacea (Testudines: Dermochelyidae): A Skeletochronological Analysis." Chelonian Conservation and Biology, 2, (2) 244–249.
• Flores-Villela, O. and Zug, George R. 1995. "Reproductive Biology of the "Chopontil," Claudius angustatus (Testudines: Kinosternidae), in Southern Veracruz, Mexico." Chelonian Conservation and Biology, 1, (3) 181–186.
• Gill, B. J., Rinke, D. R., and Zug, George R. 1995. "Emoia adspersa (Lacertilia: Scincidae)—Confirmed in Tonga." Record of Auckland Institute and Museum, 31 215–217.
• Zug, George R. 1995. "What Is a Reptile." Sci Books Films, 31, (6) 179.
• Zug, George R., Balazs, George H., and Wetherall, J. A. 1995. "Growth in Juvenile Loggerhead Seaturtles (Caretta-Caretta) in the North Pacific Pelagic Habitat." Copeia, (2) 484–487.
• Zug, George R. and Ineich, I. 1995. "A New Skink (Emoia, Lacertilia, Reptilia) from the Forest of Fiji." Proceedings of the Biological Society of Washington, 108, (3) 395–400.
• Zug, George R. and Moon, B. R. 1995. "Systematics of the Pacific Slender-Toed Geckos, Nactus-Pelagicus Complex - Oceania, Vanuatu, and Solomon-Islands Populations." Herpetologica, 51, (1) 77–90.
• Crombie, Ronald I. and Zug, George R. 1994. "kempii is the Correct Spelling for Kemp's Ridley Turtle." Marine Turtle Newsletter, (67) 2.
• Ernst, C. H. and Zug, George R. 1994. "Observations on the Reproductive-Biology of the Spotted Turtle, Clemmys-Guttata, in Southeastern Pennsylvania." Journal of Herpetology, 28, (1) 99–102.
• Ineich, I. and Zug, George R. 1994. "Un Lézard Endémique des Iles Fidji, le Gecko de Mann, Lepidodactylus manni Schmidt, 1923." Revue francaise d'Aquariologie, 20 93–96.
• Zug, George R. 1994. "[Review of] Snakes." A Natural History.Science Books & Films, 30, (6) 167.
• Zug, George R. 1994. "North American box turtles. A continuing concern." Newsletter of the Virginia Hepetological Society, 4, (2) 1–2.
• Zug, George R. 1994. "[Review of] Endoglyphs and Other Major Venomous Snakes of the World. A Checklist. 1993. By P. Golay, et al." Copeia, 1994, (4) 1069–1071.
• Zug, George R. and Ernst, C. H. 1994. "Lepidochelys." Catalogue of American Amphibians and Reptiles, 587 1–6.
• Zug, George R. and Ineich, Ivan. 1993. "Review of the biology and morphology of the Fijian bola, Ogmodon vitianus (Elapidae)." The Snake, 25 9–20.
• Ineich, I. and Zug, George R. 1991. "Nomenclatural Status of Emoia-Cyanura (Lacertilia, Scincidae) Populations in the Central Pacific." Copeia, (4) 1132–1136.
• Ineich, Ivan and Zug, George R. 1991. "Nomenclatural Status of Emoia cyanura (Lacertilia, Scincidae) Populations in the Central Pacific." Copeia, 1991, (1) 1132–1135.
• Kalb, Heather and Zug, George R. 1990. "Age Estimates for a Population of American Toads Bufo americanus (Salientia: Bufonidae)), in Northern Virginia." Brimleyana, 16 79–86.
• Zug, George R. 1990. "Review of: Reptiles and Amphibians." Copeia, 599.
• Zug, George R. 1990. "Age Determination of Long-lived Reptiles: Some Techniques for Seaturtles." Annales des Sciences Naturelles, Zoologie, Paris, 11 219–222.
• Mitchell, Joseph C. and Zug, George R. 1989. "Ecological Observations on the Gecko Hemidactylus brookii in Nepal." Amphibia-Reptilia, 9 405–413.
• Zug, George R. 1989. "Review of: Living Snakes of the World." American Scientist, 77 594–595.
• Zug, George R. 1989. "Heteronota pelagica Girard, 1857 (currently Gymnodactylus, Cyrtodactylus, Nactus pelagicus; Reptilia, Sauria): Proposed Conservation of the Specific Name." The Bulletin of Zoological Nomenclature, 46 38–40.
• Zug, George R. and Kalb, Heather. 1989. "Skeletochronological Age Estimates for Juvenile Lepidochelys kempii from the Atlantic Coast of North America." NOAA Tech.Memo, 271–273.
• Zug, George R., Springer, Victor G., Williams, Jeffrey T., and Johnson, G. David. 1989. "The Vertebrates of Rotuma and Surrounding Waters." Atoll Research Bulletin, 316 1–25. https://doi.org/10.5479/si.00775630.316.1.
• Zug, George R. 1988. "Frogs and Toads of the World." Science Activities, 25, (4) 44.
• Zug, George R. 1988. "Biological Biography of Philippine Butaan. Gray's Monitor Lizard." Naturalist Review, 6–7.
• Zug, George R. 1987. "Amphibians and Reptiles of the Outamba-Kilimi Region, Sierra Leone." Journal of the Herpetological Association of Africa, 33 1–4.
• Zug, George R. and Rand, Austin Stanley. 1987. "Estimation of Age in Nesting Female Iguana iguana: Testing Skeletochronology in a Tropical Lizard." Amphibia-Reptilia, 8 237–250.
• Zug, George R. 1986. "Sternotherus." Catalogue of American Amphibians and Reptiles, 397 1–3.
• Wynn, Addison H. and Zug, George R. 1985. "Observations on the Reproductive Biology of Candoia carinata (Serpentes, Boidae)." The Snake, 17 15–24.
• Zug, George R. 1985. "A New Skink (Reptilia, Sauria, Leiolopisma) from Fiji." Proceedings of the Biological Society of Washington, 98, (1) 221–231.
• Zug, George R. 1985. "Cyrtodactylus pelagicus: Correct Usage." Herpetologica, 16, (3) 67.
• Zug, George R. 1985. "What's Under That Rock?" Science Activities, 22, (4) 45.
• Zug, George R. 1985. "Pacific Island Lizards - Status of Type Specimens from the United-States Exploring Expedition 1838-1842." Copeia, (1) 150–154.
• Zug, George R. 1985. "Review of "The Rattlesnake"." Science Books & Films, 20, (4) 219.
• Zug, George R. 1985. "Anuran Locomotion - Fatigue and Jumping Performance." Herpetologica, 41, (2) 188–194.
• Zug, George R. 1985. "Review of "The Alligator"." Science Books & Films, 20, (5) 309.
• Zug, George R. and Balazs, George H. 1985. "Skeletochronological Age Estimates for Hawaiian Green Turtles." Marine Turtle Newsletter, 33 9–10.
• Mitchell, Joseph C. and Zug, George R. 1984. "Spermatogenic cycle of Nerodia taxispilota (Serpentes: Colubridae) in South Central Virginia." Herpetologia, 40, (2) 200–204.
• Zug, George R. 1984. "[Review]. "Advances in Herpetology and Evolutionary Biology. Essays in Honor of Ernest E. Williams"." Copeia, (2) 554–556.
• Zug, George R. 1983. "Natural History Notes on the Fijian Ground Frog (Ranidae: Platymantis vitianus)." Herpetological Review, 14, (3) 68–69.
• Zug, George R., Wynn, Addison H., and Ruckdeschel, Carol. 1983. "Age Estimates of Cumberland Island Loggerhead Sea Turtles." Marine Turtle News, (25) 9–11.
• Ruckdeschel, C. and Zug, George R. 1982. "Mortality of Sea Turtles Caretta-Caretta in Coastal Waters of Georgia." Biological Conservation, 22, (1) 5–9.
• Zug, George R. 1982. "Comments on the Proposed Conservation of Rana sphenocephala Cope, 1886. Z.N.(S.) 2141." The Bulletin of Zoological Nomenclature, 39, (2) 80–82.
• Zug, George R. 1982. "Pacific Herpetology: Some Recent Books." Herpetologica, 38, (3) 445–446.
• Zug, George R., Barber, M. M., and Dudley, J. C. 1982. "Gonadal Histology and Reproduction in Carlia-Bicarinata (Scincidae, Sauria, Reptilia) of the Port-Moresby Area, Papua-New-Guinea." Herpetologica, 38, (3) 418–425.
• Zug, George R. 1981. "1980 Marine Turtle Stranding Survey, Cumberland Island, Georgia." SEAN Bulletin, 6, (2) 20–21.
• Menzies, J. and Zug, George R. 1979. "Papuan Tree Frogs of the Litoria thesaurensis Group (Salientia: Hylidae)." Micronesica, 15, (1-2) 325–333.
• Zug, George R. and Altig, Ronald. 1979. "Anuran Locomotion—Structure and Function: The Jumping Forces of Frogs." Journal of the Washington Academy of Sciences, 68, (4) 144–147.
• Zug, George R. and Zug, P. B. 1979. "The marine toad, Bufo marinus: a natural history resume of native populations." Smithsoian Contributions to Zoology, 284.
• Zug, George R. 1978. "Once More into the Loch." 1978 Yearbook of Science and the Future, 154–169.
• Orejas-Moranda, B., Zug, George R., Garcia, D., and Achaval, F. 1977. "Scale Organs on the Head of Leptotyphlops (Reptiles, Serpentes): A Variational Study." Proceedings of the Biological Society of Washington, 90, (2) 209–213.
• Zug, George R. 1977. "Distribution and Variation of Leptotyphlops-Tricolor." Copeia, (4) 744–745.
• Zug, George R. 1977. "The Matamata (Testudines: Chelidae) Is Chelus not Chelys." Herpetologica, 33, (1) 53–54.
• Busack, S. D. and Zug, George R. 1976. "Observations on the Tadpoles of Pelobates cultripes from Southern Spain." Herpetologica, 32, (2) 151–160.
• Wake, D. B., Zweifel, R. G., Dessauer, H. C., Wace, G. W., Pianka, E. R., Rabb, G. B., Ruibal, R., Wright, J. W., and Zug, George R. 1975. "Report of the Committee on Resources in Herpetology." Copeia, 1975, (2) 391–404.
• Zug, George R., Lindgren, E., and Pippet, J. R. 1975. "Distribution and Ecology of Marine Toad, Bufo-Marinus, in Papua-New-Guinea." Pacific Science, 29, (1) 31–50.
• Orejàs-Miranda, Braulio R. and Zug, George R. 1974. "A New Tricolor Leptotyphlops (Reptilia: Serpentes) from Peru." Proceedings of the Biological Society of Washington, 87 167–174.
• Zug, George R. 1974. "Crocodilian Galloping - Unique Gait for Reptiles." Copeia, (2) 550–552.
• Zug, George R. 1973. "Searching for Hidden Animals. An Inquiry into Zoological Mysteries. [Review]." Cryptozoology, 1 84–85.
• Zug, George R. 1972. "Anuran Locomotion - Structure and Function .1. Preliminary Observations on Relation between Jumping and Osteometrics of Appendicular and Postaxial Skeleton." Copeia, (4) 613–624.
• Zug, George R. 1972. "Critique of Walk Pattern Analysis of Symmetrical Quadrupedal Gaits." Animal Behaviour, 20, (3) 436–438.
• Zug, George R. 1972. "Walk Pattern Analysis of Cryptodiran Turtle Gaits." Animal Behaviour, 20, (3) 439–443.
• Zug, George R. 1971. "American Musk Turtles, Sternothaerus or Sternotherus." Herpetologica, 27, (4) 446–449.
• Zug, George R. 1971. "Buoyancy, Locomotion, Morphology of the Pelvic Girdle and Hindlimbs and Systematics of Crypotodiran Turtles." Miscellaneous Publications, Museum of Zoology, University of Michigan, (142) 1–98.
• Zug, George R. and Schwartz, A. 1971. ""Deirochelys, D. reticularia."." Catalogue of American Amphibians and Reptiles, (107) 1–3.
• Zug, George R. 1970. "Intergradation of the Two Rhineura (Reptile) Populations in Central Florida and Comments on Its Scale Reduction." Journal of Herpetology, 4 123–129.
• Zug, George R. and Crombie, Ronald I. 1970. "Modifications of the Taylor Enzyme Method of Clearing and Staining for Amphibians and Reptiles." Herpetological Review, 2 49–50.

Book Chapter

• Kaiser, Hinrich, Zug, George R., and Bauer, Aaron M. 2018. "Germany’s Zoological Collections: an International and Personal View on an Important Historical and Contemporary Scientific Resource." In Zoological Collections of Germany. Beck, L. A., editor. 7–15. Springer International Publications.
• Gotte, Steve W., Jacobs, Jeremy F., and Zug, George R. 2016. "Preserving reptiles for research." In Reptile Ecology and Conservation, A Handbook of Techniques. Dodd, C. Kenneth, Jr., editor. 73–86. London: Oxford University Press. https://doi.org/10.1093/acprof:oso/9780198726135.003.0006.
• Zug, George R. 2011. "Tropical Asian Dry Forest Amphibians and Reptiles: A regional comparison of ecological communities." In The Ecology and Conservation of Seasonally Dry Forest in Asia. McShea, William J., Davies, Stuart James, and Bjumpakpjan, Naris, editors. 275–303. Washington, DC: Smithsonian Institution Scholarly Press.
• Zug, George R. 2009. "Reptiles (Vertebrata: Reptilia) of the Gulf of Mexico." In Gulf of Mexico origin, waters, and biota. Volume 1, Biodiversity. Felder, Darryl L. and Camp, David K., editors. 1317–1320. College Station, TX: Texas A&M Press.
• Zug, George R. 2004. "Foreword." In Snakes of India. A Field Guide. Whitaker, R. and Captain, A., editors. vii–viii. Draco Press.
• Zug, George R. 2001. "Turtles of the Lee Creek Mine (Pliocene: North Carolina)." In Geology and paleontology of the Lee Creek Mine, North Carolina, III. 203–218. Washington, D.C.: Smithsonian Institution Press. In Smithsonian Contributions to Paleobiology, 90. https://doi.org/10.5479/si.00810266.90.203.
• Zug, George R. 1999. "Foreword:." In Caribbean Amphibians and Reptiles. Crother, B. I., editor. xi. San Diego: Academic Press.
• Bjorndal, Karen and Zug, George R. 1995. "Growth and Age of Sea Turtles." In Biology and Conservation of Sea Turtles. Bjorndal, K. A., editor. 599–600. Washington, DC: Smithsonian Institution Press.
• Zug, George R. and Mitchell, J. C. 1995. "Amphibians and Reptiles of the Royal Chitwan National Park, Nepal." In Asiatic Herpetological Research. 172–180.
• Zug, George R. 1989. "Walking, Running, and Crawling." In Crocodiles and Alligators. Ross, C. A., editor. 46. Oxford: Facts on File, New York.
• Zug, George R. 1986. "Leaps and Bounds. Why Have Frogs Lost Their Tails?" In The Encyclopedia of Reptiles and Amphibians. Halliday, T. R. and Adler, K., editors. 56–57.
• Zug, George R. 1983. "Bufo marinus." In Costa Rican Natural History. Janzen, D., editor. 386–387.
• Zug, George R. 1979. "The Endoskeleton: The Comparative Anatomy of the Girdles, the Sternum, and the Paired Appendages." In Comparative Vertebrate Anatomy. Wake, M., editor. 238–264. Chicago: University of Chicago Press.
• Zug, George R. 1974. "Locomotion." In Encyclopaedia Britannica. 15–24.
• Zug, George R. 1971. "The Musculoskeletal System: An Evolutionary Perspective." In Chordate Structure and Function. 200–241. New York: Macmillan Co.

Book Review

• Zug, George R. 2006. [Book review] "A Sheltered Life. The Unexpected History of the Giant Tortoise." Herpetological Review, 37, (3), 374.
• Zug, George R. 1985. [Book review] ""The Book of Indian Reptiles" by J. C. Daniel." Herpetologica, 41, (2), 236–237.