Michael Carleton

• Ph.D. (Zoology): University of Michigan, 1979
• B.S. (Wildlife Biology): University of Massachusetts, 1966

Systematics and evolution of muroid rodents.

Journal Article

• Pradhan, Nelish, Norris, Ryan W., Decher, Jan, Kerbis Peterhans, Julian, Gray, Christopher R., Bauer, George, Carleton, Michael D., and Kilpatrick, C. William. 2021. "Phylogenetic relationships and biogeography of the Hybomys division (Muridae: Murinae: Arvicanthini), rodents endemic to Africa's rainforests." Journal of Vertebrate Biology, 70, (2). https://doi.org/10.25225/jvb.21034.
• Jansa, Sharon A., Carleton, Michael D., Soarimalala, Voahangy, Rakotomalala, Zafimahery, and Goodman, Steven M. 2019. "A Review of the Eliurus Tanala Complex (Rodentia: Muroidea: Nesomyidae), with Description of a New Species from Dry Forests of Western Madagascar." Bulletin of the American Museum of Natural History, (430) 1–67. https://doi.org/10.1206/0003-0090.430.1.1.
• Carleton, Michael D., Banasiak, Rebecca A., and Stanley, William T. 2015. "A new species of the rodent genus Hylomyscus from Angola, with a distributional summary of the H. anselli species group (Muridae: Murinae: Praomyini)." Zootaxa, 4040, (2) 101–128. https://doi.org/10.11646/zootaxa.4040.2.1.
• Carleton, Michael D., Gardner, Alfred L., Pavlinov, Igor Ya, and Musser, Guy G. 2014. "The valid generic name for red-backed voles (Muroidea: Cricetidae: Arvicolinae): restatement of the case for Myodes Pallas, 1811." Journal of mammalogy, 95, (5) 943–959. https://doi.org/10.1644/14-MAMM-A-004.
• Carleton, Michael D., Smeenk, Chris, Angermann, Renate, and Goodman, Steven M. 2014. "Taxonomy of nesomyine rodents (Muroidea: Nesomyidae: Nesomyinae): Designation of lectotypes and restriction of type localities for species-group taxa in the genus Nesomys Peters." Proceedings of the Biological Society of Washington, 126, (4) 414–455. https://doi.org/10.2988/0006-324X-126.4.414.
• Carleton, Michael D. and Stanley, William T. 2012. "Species limits within the Praomys delectorum group (Rodentia: Muridae: Murinae) of East Africa: a morphometric reassessment and biogeographical implications." Zoological Journal of the Linnean Society, 165, (2) 420–469. https://doi.org/10.1111/j.1096-3642.2012.00817.x.
• Taylor, Peter J., Lavrenchenko, Leonid A., Carleton, Michael D., Verheyen, Erik, Bennett, Nigel C., Oosthuizen, Carel J., and Maree, Sarita. 2011. "Specific limits and emerging diversity patterns in East African populations of laminate-toothed rats, genus Otomys (Muridae: Murinae: Otomyini): Revision of the Otomys typus complex." Zootaxa, (3024) 1–66.
• Ding, X., Leigh, C. M., Goodman, S. M., Bedford, J. M., Carleton, Michael D., and Breed, W. G. 2010. "Sperm morphology in the Malagasy rodents (Muroidea: Nesomyinae)." Journal of Morphology, 271, (12) 1493–1500. https://doi.org/10.1002/jmor.10889.
• Carleton, Michael D. 2009. "They sort out like nuts and bolts: a scientific biography of Guy G. Musser." Bulletin of the American Museum of Natural History, 331 4–32.
• Carleton, Michael D. and Arroyo-Cabrales, Joaquin. 2009. "Review of the Oryzomys couesi complex (Rodentia: Cricetidae: Sigmodontinae) in western Mexico." Bulletin of the American Museum of Natural History, 331 94–127.
• Carleton, Michael D., Emmons, Louise H., and Musser, Guy G. 2009. "A new species of the rodent genus Oecomys (Cricetidae: Sigmodontinae: Oryzomyini) from eastern Bolivia, with emended definitions of O. concolor (Wagner) and O. mamorae (Thomas)." American Museum Novitates, 3661 1–32.
• Gardner, Alfred L. and Carleton, Michael D. 2009. "A new species of Reithrodontomys, subgenus Aporodon (Cricetidae: Neotominae), from the highlands of Costa Rica, with comments on Costa Rican and Panamanian Reithrodontomys." Bulletin of the American Museum of Natural History, 331 157–182.
• Carleton, Michael D. and Goodman, S. M. 2007. "A new species of the Eliurus majori complex (Rodentia: Muroidea: Nesomyidae) from south-central Madagascar, with remarks on emergent species groupings in the genus Eliurus." American Museum Novitates, 3547 21.
• Carleton, Michael D. and Byrne, E. Schaefer. 2006. "The status of Otomys orestes dollmani Heller, 1912 (Muridae: Otomyinae), a rodent described from the Mathews Range, central Kenya." Proceedings of the Biological Society of Washington, 119, (4) 477–515. https://doi.org/10.2988/0006-324X(2006)119[477:TSOOOD]2.0.CO;2.
• Carleton, Michael D., Kerbis Peterhans, J. C., and Stanley, W. T. 2006. "Review of the Hylomyscus denniae group (Rodentia: Muridae) in eastern Africa, with comments on the generic allocation of Epimys endorobae Heller." Proceedings of the Biological Society of Washington, 119, (2) 293–325. https://doi.org/10.2988/0006-324X(2006)119[293:ROTHDG]2.0.CO;2.
• O'Brien, C., McShea, William J., Guimondou, S., Barriere, P., and Carleton, Michael D. 2006. "Terrestrial small mammals (Soricidae and Muridae) from the Gamba Complex in Gabon: species composition and comparison of sampling tecnhiques." Bulletin of the Biological Society of Washington, 12 353–363.
• Carleton, Michael D. and Lawlor, T. E. 2005. "Peromyscus from Santa Catalina Island, Sea of Cortez, Mexico: taxonomic identities and biogeographic implications." Journal of Mammalogy, 86, (4) 814–825.
• Carleton, Michael D. and Stanley, W. T. 2005. "Review of the Hylomyscus denniae complex (Rodentia: Muridae) in Tanzania, with a description of a new species." Proceedings of the Biological Society of Washington, 118, (3) 619–646.
• Jenkins, P. D. and Carleton, Michael D. 2005. "Charles Immanuel Forsyth Major's expedition to Madagascar, 1894 to 1896: beginnings of modern systematic study of the island's mammalian fauna." Journal of Natural History, 39, (20) 1779–1818.
• Carleton, Michael D., Sanchez, O., and Vidales, G. Urbano. 2002. "A New Species of Habromys (Muridae: Neotominae) from Mexico, with Generic Review of Species Definitions and Remarks on Diversity Patterns Among Mesoamerican Small Mammals Restricted to Humid Montane Forests." Proceedings of the Biological Society of Washington, 115, (3) 488–533.
• Carleton, Michael D., Goodman, S. M., and Rakotondravony, D. 2001. "A new species of tufted-tailed rat, genus Eliurus (Muridae : Nesomyinae), from western Madagascar, with notes on the distribution of E-myoxinus." Proceedings of the Biological Society of Washington, 114, (4) 972–987.
• Carleton, Michael D. and Goodman, S. M. 2000. "Rodents of the Parc National de Marojejy, Madagascar." Fieldiana. Zoology, 97 231–263.
• Carleton, Michael D., Fisher, Robert D., and Gardner, Alfred L. 1999. "Identification and distribution of cotton rats, genus Sigmodon (Muridae : Sigmodontinae), of Nayarit, Mexico." Proceedings of the Biological Society of Washington, 112, (4) 813–856.
• Carleton, Michael D. and Olson, Storrs L. 1999. "Amerigo Vespucci and the Rat of Fernando de Noronha: a New Genus and Species of Rodentia (Muridae: Sigmodontinae) from a Volcanic Island off Brazil's Continental Shelf." American Museum Novitates, 3256 1–59.
• Goodman, S. M., Carleton, Michael D., and Pidgeon, M. 1999. "The Rodents of the Reserve Naturelle Integrale d'Andohahela, Madagascar." Fieldiana. Zoology, 94 217–249.
• Carleton, Michael D. and Goodman, S. M. 1998. "New Taxa of Nesomyine Rodents (Muroidea: Muridae) from Madagascar's Northern Highlands, with Taxonomic Comments on Previously Described Forms." Fieldiana. Zoology, 90 163–200.
• Goodman, S. M. and Carleton, Michael D. 1998. "The Rodents of the Reserve Speciale d'Anjanaharibe-Sud, Madagascar." Fieldiana. Zoology, 90 201–221.
• Musser, G. G., Carleton, Michael D., Brothers, E. M., and Gardner, Alfred L. 1998. "Systematic studies of oryzomyine rodents (Muridae, Sigmodontinae): Diagnoses and distributions of species formerly assigned to Oryzomys "capito"." Bulletin of the American Museum of Natural History, (236) 10–+.
• Carleton, Michael D. and Van der Straeten, E. 1997. "Morphological differentiation among Subsaharan and north African populations of the Lemniscomys barbarus complex (Rodentia : Muridae)." Proceedings of the Biological Society of Washington, 110, (4) 640–680.
• Carleton, Michael D. and Goodman, S. M. 1996. "The rodents of the Réserve Naturelle Intégrale d'Andringitra, Madagascar." Fieldiana. Zoology, 85 257–283.
• Carleton, Michael D. and Goodman, S. M. 1996. "Systematic Studies of Madagascar's Endemic Rodents (Muroidea: Nesomyinae): A New Genus and Species from the Central Highlands." Fieldiana. Zoology, 85 231–256.
• Musser, G. G., Brothers, E. M., Carleton, Michael D., and Hutterer, R. 1996. "Taxonomy and Distributional Records of Oriental and European Apodemus, with a Review of the Apodemus-Sylvaemus Problem." Bonner Zoologische Beitrage, Bonn, 46, (1-4) 143–190.
• Carleton, Michael D. and Musser, G. G. 1995. "Systematic Studies of Oryzomyine Rodents (Muridae, Sigmodontinae) - Definition and Distribution of Oligoryzomys-Vegetus (Bangs, 1902)." Proceedings of the Biological Society of Washington, 108, (2) 338–369.
• Brown, J. H., Carleton, Michael D., Estes, J. A., Fleming, T. H., and Musser, G. G. 1994. "Hooper,emmet,t. - 1911-1992 - Obituary." Journal of Mammalogy, 75, (3) 787–792.
• Carleton, Michael D. 1994. "Systematic studies of Madagascar's endemic rodents (Muroidea: Nesomyinae): revision of the genus Eliurus." American Museum Novitates, 3087 1–55.
• Voss, R. S. and Carleton, Michael D. 1993. "A new genus for Hesperomys molitor Winge and Holochilus magnus Hershkovitz (Mammalia, Muridae) with an analysis of its phylogenetic relationships." American Museum Novitates, 3085 1–39.
• Carleton, Michael D. and Martinez, Christine. 1991. "Morphometric Differentiation among West African Populations of the Rodent Genus Dasymys (Muroidea, Murinae), and its Taxonomic Implications." Proceedings of the Biological Society of Washington, 104, (3) 419–435.
• Carleton, Michael D. and Musser, G. G. 1989. "Systematic Studies of Oryzomyine Rodents (Muridae, Sigmodontinae): A Synopsis of Microryzomys." Bulletin of the American Museum of Natural History, 191 1–83.
• Carleton, Michael D. and Robbins, C. B. 1985. "On the Status and Affinities of Hybomys-Planifrons (Miller, 1900) (Rodentia, Muridae)." Proceedings of the Biological Society of Washington, 98, (4) 956–1003.
• Carleton, Michael D. 1981. "A Survey of Gross Stomach Morphology in Microtinae (Rodentia, Muroidea)." Zeitschrift Fur Saugetierkunde-International Journal of Mammalian Biology, 46, (2) 93–108.
• Carleton, Michael D. 1980. "Phylogenetic Relationships in Neotomine-Peromyscine Rodents (Muroidea) and a Reappraisal of the Dichotomy within New World Cricetinae." Miscellaneous Publications Museum of Zoology, University of Michigan, 157 1–146.
• Carleton, Michael D. 1979. "Taxonomic Status and Relationships of Peromyscus boylii from El Salvador." Journal of mammalogy, 60 280–296.
• Carleton, Michael D. and Eshelman, R. E. 1979. "A Synopsis of Fossil Grasshopper Mice, Genus Onychomys, and Their Relationships to Recent Species." Papers on Paleontology, University of Michigan Museum of Paleontology, 21 1–63.
• Carleton, Michael D. and Myers, P. 1979. "Karyotypes of Some Harvest Mice, Genus Reithrodontomys." Journal of mammalogy, 60 307–313.

Book Chapter

• Carleton, Michael D. 2015. "Genus Microryzomys Thomas, 1917." In Mammals of South America, Volume 2. Rodents. Patton, James L., Pardinas, Ulyses F. J., and D'Elia, Guillermo, editors. 355–359. Chicago: The University of Chicago Press.
• Carleton, Michael D. 2015. "Genus Transandinomys Weksler, Percequillo, and Voss, 2006." In Mammals of South America, Volume 2. Rodents. Patton, James L., Pardinas, Ulyses F. J., and D'Elia, Guillermo, editors. 456–460. Chicago: The University of Chicago Press.
• Carleton, Michael D. and Musser, Guy G. 2015. "Genus Oecomys Thomas, 1906." In Mammals of South America, Volume 2. Rodents. Patton, James L., Pardinas, Ulyses F. J., and D'Elia, Guillermo, editors. 393–417. Chicago: The University of Chicago Press.
• Carleton, Michael D. 2013. "Genus Hybomys Forest Mice." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 420–421. London: Bloomsbury Publishing.
• Carleton, Michael D. 2013. "Hybomys planifrons Liberian Forest Mouse." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 424–425. London: Bloomsbury Publishing.
• Carleton, Michael D. 2013. "Dasymys nudipes Angolan Shaggy Rat." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 397–398. London: Bloomsbury Publishing.
• Carleton, Michael D. 2013. "Dasymys foxi Fox's Shaggy Rat." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 394–395. London: Bloomsbury Publishing.
• Carleton, Michael D. 2013. "Thallomys loringi Loring's Acacia Rat." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 558–559. London: Bloomsbury Publishing.
• Carleton, Michael D. 2013. "Genus Dasymys Shaggy Rats." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 392–394. London: Bloomsbury Publishing.
• Carleton, Michael D. 2013. "Dasymys montanus Montane Shaggy Rat." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 396–397. London: Bloomsbury Publishing.
• Carleton, Michael D. 2013. "Genus Lemniscomys Grass Mice." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 441–443. London: Bloomsbury Publishing.
• Carleton, Michael D. 2013. "Subfamily Otomyinae - Vlei Rats and Whistling Rats." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 571–573. London: Bloomsbury Publishing.
• Carleton, Michael D. and Happold, David C. D. 2013. "Order Rodentia - Rodents." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 27–37. London: Bloomsbury Publishing.
• Carleton, Michael D. and Happold, David C. D. 2013. "Hybomys trivirgatus Three-striped Forest Mouse." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 425–426. London: Bloomsbury Publlishing.
• Carleton, Michael D. and Musser, Guy G. 2013. "Family Cricetidae." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 210. London: Bloomsbury Publishing.
• Carleton, Michael D. and Musser, Guy G. 2013. "Subfamily Arvicolinae - Voles, Lemmings and Muskrats." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 210–211. London: Bloomsbury Publishing.
• Carleton, Michael D. and Musser, Guy G. 2013. "Subfamily Gerbillinae - Gerbils and Jirds." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 260–262. London: Bloomsbury Publishing.
• Carleton, Michael D. and Musser, Guy G. 2013. "Subfamily Lophiomyinae - Maned Rat." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 212–213. London: Bloomsbury Publishing.
• Carleton, Michael D. and Musser, Guy G. 2013. "Family Nesomyidae." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 153. London: Bloomsbury Publishing.
• Carleton, Michael D. and Musser, Guy G. 2013. "Subfamily Cricetomyinae - Pouched Rats and Pouched Mice." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 153–154. London: Bloomsbury Publishing.
• Carleton, Michael D. and Musser, Guy G. 2013. "Family Muridae." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 216. London: Bloomsbury Publishing.
• Carleton, Michael D. and Musser, Guy G. 2013. "Subfamily Petromyscinae - Pigmy Rock Mice." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 203–204. London: Bloomsbury Publishing.
• Carleton, Michael D. and Musser, Guy G. 2013. "Subfamily Mystromyinae - White-tailed Rat." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 201. London: Bloomsbury Publishing.
• Carleton, Michael D. and Musser, Guy G. 2013. "Subfamily Deomyinae - Spiny Mice, Link Rat and Brush-furred Rats." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 217. London: Bloomsbury Publishing.
• Musser, Guy G. and Carleton, Michael D. 2013. "Subfamily Leimacomyinae - Buttner's Forest Mouse." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 358–359. London: Bloomsbury Publishing.
• Musser, Guy G. and Carleton, Michael D. 2013. "Family Spalacidae." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits.. Happold D. C. D., editor. 143–144. London: Bloomsbury Publishing.
• Musser, Guy G. and Carleton, Michael D. 2013. "Subfamily Murinae - Rats and Mice." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 361–362. London: Bloomsbury Publishing.
• Musser, Guy G. and Carleton, Michael D. 2013. "Subfamily Tachyoryctincae - African Root-rats." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits.. Happold, D. C. D., editor. 147–148. London: Bloomsbury Publishing.
• Musser, Guy G. and Carleton, Michael D. 2013. "Subfamily Spalacidae - Mole Rats." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 144–148. London: Bloomsbury Publishing.
• Musser, Guy G. and Carleton, Michael D. 2013. "Subfamily Delanymyinae - Delany's Swamp Mouse." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 165–166. London: Bloomsbury Publishing.
• Musser, Guy G. and Carleton, Michael D. 2013. "Subfamily Dendromurinae - African Climbing Mice." In Mammals of Africa, Volume III: Rodents, Hares and Rabbits. Happold, David C. D., editor. 168–169. London: Bloomsbury Publishing.
• Carleton, Michael D. and Musser, Guy G. 2005. "Order Rodentia." In Mammal Species of the World, Third Edition. Wilson, Don E. and Reeder, D. M., editors. 745–752. The Johns Hopkins University Press.
• Musser, Guy G. and Carleton, Michael D. 2005. "Superfamily Muroidea." In Mammal Species of the World, Third Edition. Wilson, Don E. and Reeder, D. M., editors. 894–1531. The Johns Hopkins University Press.
• Carleton, Michael D. 2003. "Eliurus, Tufted-tailed Rats." In The Natural History of Madagascar. Goodman, S. M. and Benstead, J. P., editors. 1373–1380. The University of Chicago Press.
• Carleton, Michael D. and Goodman, S. M. 2003. "Gymnuromys, Voalavoanala." In The Natural History of Madagascar. Goodman, S. M. and Benstead, J. P., editors. 1381–1383. The University of Chicago Press.
• Carleton, Michael D. and Goodman, S. M. 2003. "Rodentia: Brachytarsomys, White-tailed tree rats, Antsangy." In The Natural History of Madagascar. Goodman, S. M. and Benstead, J. P., editors. 1368–1370. The University of Chicago Press.
• Carleton, Michael D. and Goodman, S. M. 2003. "Macrotarsomys, Big-footed Mice." In The Natural History of Madagascar. Goodman, S. M. and Benstead, J. P., editors. 1386–1388. The University of Chicago Press.
• Carleton, Michael D., Musser, Guy G., and Pavlinov, I. Ya. 2003. "Myodes Pallas, 1811, is the valid name for the genus of red-backed voles." In Sytematics, Phylogeny and Paleontology of Small Mammals-- An International Conference Devoted to the 90th Anniversary of Prof. I. M. Gromov. Averianov, A. O. and Abramson, N. I., editors. 96–98. Proceedings of the Zoological Institute.
• Jansa, S. A. and Carleton, Michael D. 2003. "Systematics and phylogenetics of Madagascar's native rodents." In The Natural History of Madagascar. Goodman, S. M. and Benstead, J. P., editors. 1257–1265. The University of Chicago Press.
• Jansa, S. A. and Carleton, Michael D. 2003. "Brachyuromys, Short-tailed Rats, Malagasy Voles." In The Natural History of Madagascar. Goodman, S. M. and Benstead, J. P., editors. 1370–1373. The University of Chicago Press.
• Carleton, Michael D. 1989. "Systematics and Evolution." In Advances in the Study of Peromyscus (Rodentia). Kirkland, D. L. and Layne, J., editors. 367. Lubbock: Texas Tech University Press.
• Carleton, Michael D. 1985. "Macroanatomy." In Biology of New World Microtus. Tamarin, R. H., editor. 116–175.
• Carleton, Michael D. 1984. "Chapter 9: Introduction to Rodents." In Orders and Families of Recent Mammals of the World. Anderson, S. and Jones, J. K., editors. 255–265. New York: John Wiley & Sons.
• Carleton, Michael D. and Musser, Guy G. 1984. "Chapter 11: Muroid Rodents." In Orders and Families of Recent Mammals of the World. Anderson, S. and Jones, J. K., editors. 289–379. New York: John Wiley & Sons.
• Carleton, Michael D. and Eshelman, R. 1979. "A Synopsis of Fossil Grasshopper Mice, Genus Onychomys, and Their Relationship to Recent Species." In Contributions of Museum of Paleontology, University of Michigan. 1–63.

Book

• Voss, R. S. and Carleton, Michael D., editors. 2009. Systematic mammalogy: Contributions in honor of Guy G. Musser. New York: American Museum of Natural History. In Bulletin of the American Museum of Natural History.
• Schmidt, David F., Ludwig, Craig A., and Carleton, Michael D. 2008. The Smithsonian Institution African Mammal Project (1961-1972): An Annotated Gazetteer of Collecting Localities and Summary of Its Taxonomic and Geographic Scope. In Smithsonian Contributions to Zoology, 628. https://doi.org/10.5479/si.00810282.628.
• Carleton, Michael D. 1982. Distribution and systematics of Peromyscus (Mammalia:Rodentia) of Nayarit, Mexico. Washington: Smithsonian Institution. In Smithsonian Contributions to Zoology, 352. https://doi.org/10.5479/si.00810282.352.
• Myers, Philip and Carleton, Michael D. 1981. The Species of Oryzomys (Oligoryzomys) in Paraguay: And the Identity of Azara's "Rat Sixième Ou Rat À Tarse Noir". Museum of Zoology, University of Michigan.