Smithsonian National Museum of Natural History

Matthew Carrano

Research Geologist and Curator of Dinosauria
Department / Division
Education
  • B.S., Brown University
  • M.S., University of Chicago
  • Ph.D., University of Chicago
Research Interests

Large-scale evolutionary patterns within Dinosauria; systematics of basal Theropoda; vertebrate paleoecology of Mesozoic terrestrial ecosystems; the dinosaur fossil record.

Publications
  • Montoya-Sanhueza, Germ, Moreno, Karen, Bobe, René, Carrano, Matthew T., García, Marcelo, and Corgne, Alexandre. 2017. "Peltephilidae and Mesotheriidae (Mammalia) from late Miocene strata of Northern Chilean Andes, Caragua." Journal of South American Earth Sciences 75:51-65. https://doi.org/10.1016/j.jsames.2017.01.009
  • Benson, Roger B. J., Butler, Richard J., Alroy, John, Mannion, Philip D., Carrano, Matthew T., and Lloyd, Graeme T. 2016. "Near-Stasis in the Long-Term Diversification of Mesozoic Tetrapods." PLoS Biology 14 (1):1-27. https://doi.org/10.1371/journal.pbio.1002359
  • Carrano, Matthew T. 2016. "Vertebrate microfossil bonebeds as benchmarks for terrestrial vertebrate diversity and paleoecology through time." Given at Department of Integrated Biology, University of California, Berkeley, 2016.
  • Carrano, Matthew T. and Choiniere, Jonah. 2016. "New Information on the Forearm and Manus of Ceratosaurus Nasicornis Marsh, 1884 (Dinosauria, Theropoda), with Implications for Theropod Forelimb Evolution." Journal of Vertebrate Paleontology 36 (2):https://doi.org/10.1080/02724634.2015.1054497
  • Carrano, Matthew T., Oreska, Matthew P. J., and Lockwood, Rowan. 2016. "Vertebrate Paleontology of the Cloverly Formation (Lower Cretaceous), Ii: Paleoecology." Journal of Vertebrate Paleontology 36 (2):https://doi.org/10.1080/02724634.2015.1071265
  • Rauhut, Oliver W. M. and Carrano, Matthew T. 2016. "The theropod dinosaur Elaphrosaurus bambergi Janensch, 1920, from the Late Jurassic of Tendaguru, Tanzania." Zoological Journal of the Linnean Society 178 (3):546-610. https://doi.org/10.1111/zoj.12425
  • Aiello, B. R., Iriarte-Diaz, J., Blob, R. W., Butcher, M. T., Carrano, Matthew T., Espinoza, N. R., Main, R. P., and Ross, C. F. 2015. "Bone strain magnitude is correlated with bone strain rate in tetrapods: implications for models of mechanotransduction." Proceedings of the Royal Society B: Biological Sciences 282 (1810):https://doi.org/10.1098/rspb.2015.0321
  • Carrano, Matthew T. 2015. "A curatorial perspective on the Deep Time exhibit renovation." Given at The Future of Museum and Gallery Design, Hong Kong University, 2015.
  • Carrano, Matthew T. and D'Emic, Michael D. 2015. "Osteoderms of the titanosaur sauropod dinosaur Alamosaurus sanjuanensis Gilmore, 1922." Journal of Vertebrate Paleontology 35 (1):https://doi.org/10.1080/02724634.2014.901334
  • Evans, David C., Barrett, Paul M., Brink, Kirstin S., and Carrano, Matthew T. 2015. "Osteology and bone microstructure of new, small theropod dinosaur material from the early Late Cretaceous of Morocco." Gondwana Research 27 (3):1034-1041. https://doi.org/10.1016/j.gr.2014.03.016
  • Ksepka, Daniel T., Parham, James F., Allman, James F., Benton, Michael J., Carrano, Matthew T., Cranston, Karen A., Donoghue, Philip C. J., Head, Jason J., Hermsen, Elizabeth J., Irmis, Randall B., Joyce, Walter G., Kohli, Manpreet, Lamm, Kristin S., Leehr, Dan, Patané, José S. L., Polly, P. David, Phillips, Matthew J., Smith, N. Adam, Smith, Nathan D., van Tuinen, Marcel, Ware, Jessica L., and Warnock, Rachel C. M. 2015. "The Fossil Calibration Database, A New Resource for Divergence Dating." Systematic Biology 64 (5):853-859. https://doi.org/10.1093/sysbio/syv025
  • Mannion, Philip D., Benson, Roger B. J., Carrano, Matthew T., Tennant, Jonathan P., Judd, Jack, and Butler, Richard J. 2015. "Climate constrains the evolutionary history and biodiversity of crocodylians." Nature Communications 6 (8438):https://doi.org/10.1038/ncomms9438
  • Benson, Roger B. J., Campione, Nicolás E., Carrano, Matthew T., Mannion, Philip D., Sullivan, Corwin, Upchurch, Paul, and Evans, David C. 2014. "Rates of Dinosaur Body Mass Evolution Indicate 170 Million Years of Sustained Ecological Innovation on the Avian Stem Lineage." PLoS Biology 12 (5):1-11. https://doi.org/10.1371/journal.pbio.1001853
  • Brusatte, Stephen L., Butler, Richard J., Barrett, Paul M., Carrano, Matthew T., Evans, David C., Lloyd, Graeme T., Mannion, Philip D., Norell, Mark A., Peppe, Daniel J., Upchurch, Paul, and Williamson, Thomas E. 2014. "The extinction of the dinosaurs." Biological Reviews 90 (2):628-642. https://doi.org/10.1111/brv.12128
  • Campione, Nicolas E., Evans, David C., Brown, Caleb M., and Carrano, Matthew T. 2014. "Body mass estimation in non-avian bipeds using a theoretical conversion to quadruped stylopodial proportions." Methods in Ecology and Evolution 5 (9):913-923. https://doi.org/10.1111/2041-210X.12226
  • Carrano, Matthew T. 2014. "Challenges and benefits of using vertebrate microfossil bonebeds for understanding terrestrial paleoecosystems." Given at 4th International Palaeontological Congress, 2014.
  • Conrad, Jack L., Head, Jason J., and Carrano, Matthew T. 2014. "Unusual Soft-Tissue Preservation of a Crocodile Lizard (Squamata, Shinisauria) From the Green River Formation (Eocene) and Shinisaur Relationships." Anatomical Record : Advances in Integrative Anatomy and Evolutionary Biology 297 (3):545-559. https://doi.org/10.1002/ar.22868
  • Thomas, Daniel B., McGraw, Kevin J., Butler, Michael W., Carrano, Matthew T., Madden, Odile, and James, Helen F. 2014. "Ancient origins and multiple appearances of carotenoid-pigmented feathers in birds." Proceedings of the Royal Society B: Biological Sciences 281 (1788):https://doi.org/10.1098/rspb.2014.0806
  • Oreska, Matthew P. J., Carrano, Matthew T., and Dzikiewicz, Katherine M. 2013. "Vertebrate Paleontology of the Cloverly Formation (Lower Cretaceous), I: Faunal Composition, Biogeographic Relationships, and Sampling." Journal of Vertebrate Paleontology 33 (2):264-292. https://doi.org/10.1080/02724634.2012.717567
  • Uhen, Mark D., Barnosky, Anthony D., Bills, Brian, Blois, Jessica, Carrano, Matthew T., Carrasco, Marc A., Erickson, Gregory M., Eronen, Jussi T., Fortelius, Mikael, Graham, Russell W., Grimm, Eric C., O'Leary, Maureen A., Mast, Austin, Piel, William H., Polly, P. David, and Saila, Laura K. 2013. "From Card Catalogs to Computers: Databases in Vertebrate Paleontology." Journal of Vertebrate Paleontology 33 (1):13-28. https://doi.org/10.1080/02724634.2012.716114
  • Benson, Roger B. J., Butler, Richard J., Carrano, Matthew T., and O'Connor, Patrick M. 2012. "Air-filled postcranial bones in theropod dinosaurs: physiological implications and the 'reptile'–bird transition." Biological Reviews 87 (1):168-193. https://doi.org/10.1111/j.1469-185X.2011.00190.x
  • Burch, Sara H. and Carrano, Matthew T. 2012. "An Articulated Pectoral Girdle and Forelimb of the Abelisaurid Theropod Majungasaurus Crenatissimus from the Late Cretaceous of Madagascar." Journal of Vertebrate Paleontology 32 (1):1-16. https://doi.org/10.1080/02724634.2012.622027
  • Carrano, Matthew T. 2012. "Paleoecology and biogeography of the vertebrate fauna of the Morrison Formation, Late Jurassic, western United States." Given at Tendaguru and Beyond : Vertebrates in Jurassic Ecosystems (Symposium), Centenary Meeting, Paläontologische Gessellschaft, 2012.
  • Carrano, Matthew T. 2012. "Dinosaur detective stories: what do we know about dinosaurs, and how do we know it?." Given at Undergraduate Research and Design Symposium. Northern Arizona University., 2012.
  • Carrano, Matthew T., Benson, Roger B. J., and Sampson, Scott D. 2012. "The phylogeny of Tetanurae (Dinosauria: Theropoda)." Journal of Systematic Palaeontology 10 (2):211-300. https://doi.org/10.1080/14772019.2011.630927
  • Fitzgerald, Erich M. G., Carrano, Matthew T., Holland, Timothy, Wagstaff, Barbara E., Pickering, David, Rich, Thomas H., and Vickers-Rich, Patricia. 2012. "First ceratosaurian dinosaur from Australia." Naturwissenschaften 99 (5):397-405. https://doi.org/10.1007/s00114-012-0915-3
  • Thomas, Daniel B., James, Helen F., Carrano, Matthew T., and Madden, Odile. 2012. "Searching for Evidence of Fossil Feather Color with Spectroscopy." Journal of Vertebrate Paleontology 32:183-184.
  • Butler, Richard J., Benson, Roger B. J., Carrano, Matthew T., Mannion, Philip D., and Upchurch, Paul. 2011. "Sea level, dinosaur diversity and sampling biases: investigating the ‘common cause’ hypothesis in the terrestrial realm." Proceedings of the Royal Society B: Biological Sciences 278 (1709):1165-1170. https://doi.org/10.1098/rspb.2010.1754
  • Carrano, Matthew T. 2011. "The evolutionary history of basal tetanuran theropod dinosaurs." Given at IV Congreso Latinoamericano Paleontología de Vertebrados, .
  • Carrano, Matthew T., Loewen, Mark A., and Sertich, Joseph J. W. 2011. New materials of Masiakasaurus knopfleri Sampson, Carrano, and Forster, 2001, and implications for the morphology of the Noasauridae (Theropoda:Ceratosauria). Smithsonian Institution Scholarly Press. https://doi.org/10.5479/si.00810266.95.1
  • Hall, Margaret I., Kirk, E. C., Kamilar, Jason M., and Carrano, Matthew T. 2011. "Comment on "Nocturnality in Dinosaurs Inferred from Scleral Ring and Orbit Morphology"." Science 334 (6063):1641-1641. https://doi.org/10.1126/science.1208442
  • Ksepka, Daniel T., Benton, Michael J., Carrano, Matthew T., Gandolfo, Maria A., Head, Jason J., Hermsen, Elizabeth J., Joyce, Walter G., Lamm, Kristin S., Patané, José S. L., Phillips, Matthew J., Polly, P. David, van Tuinen, Marcel, Ware, Jessica L., Warnock, Rachel C. M., and Parham, James F. 2011. "Synthesizing and databasing fossil calibrations: divergence dating and beyond." Biology Letters 7 (6):801-803. https://doi.org/10.1098/rsbl.2011.0356
  • Mannion, Philip D., Benson, Roger B. J., Upchurch, Paul, Butler, Richard J., Carrano, Matthew T., and Barrett, Paul M. 2011. "A temperate palaeodiversity peak in Mesozoic dinosaurs and evidence for Late Cretaceous geographical partitioning." Global Ecology and Biogeography 21 (9):898-908. https://doi.org/10.1111/j.1466-8238.2011.00735.x
  • Mannion, Philip D., Upchurch, Paul, Carrano, Matthew T., and Barrett, Paul M. 2011. "Testing the effect of the rock record on diversity: a multidisciplinary approach to elucidating the generic richness of sauropodomorph dinosaurs through time." Biological Reviews 86 (1):157-181. https://doi.org/10.1111/j.1469-185X.2010.00139.x
  • Upchurch, P., Mannion, P. D., Benson, R. B. J., Butler, R. J., and Carrano, Matthew T. 2011. "Geological and anthropogenic controls on the sampling of the terrestrial fossil record: a case study from the Dinosauria." Geological Society, London, Special Publications 358 (1):209-240. https://doi.org/10.1144/SP358.14
  • Wilson, Jeffrey A., Barrett, Paul M., and Carrano, Matthew T. 2011. "An associated partial skeleton of Jainosaurus cf. septentrionalis (Dinosauria: Sauropoda) from the Late Cretaceous of Chhota Simla, Central India." Palaeontology 54 (5):981-998. https://doi.org/10.1111/j.1475-4983.2011.01087.x
  • Benson, Roger B. J., Carrano, Matthew T., and Brusatte, Stephen L. 2010. "A new clade of archaic large-bodied predatory dinosaurs (Theropoda: Allosauroidea) that survived to the latest Mesozoic." Naturwissenschaften 97 (1):71-78. https://doi.org/10.1007/s00114-009-0614-x
  • Carrano, Matthew T., Wilson, Jeffrey A., and Barrett, Paul M. 2010. "The history of dinosaur collecting in central India, 1828-1947." in Dinosaurs and other Extinct Saurians: a Historical Perspective, edited by Moody, R. T. J., Buffetaut, E., Naish, D., and Martill, D. M., 161-173. Geological Society.
  • Hunt, Gene and Carrano, Matthew T. 2010. "Models and methods for analyzing phenotypic evolution in lineages and clades." in Quantitative Methods in Paleobiology, edited by Alroy, John and Hunt, Gene., 245-269. The Paleontological Society.
  • Carrano, Matthew T. 2009. "The phylogeny of megalosauroids (Dinosauria: Theropoda) with implications for the evolution of North African ecosystems." Given at First International Congress on North African Vertebrate Palaeontology, 2009.
  • Carrano, Matthew T., Krause, D. W., O'Connor, P. M., and Sampson, S. D. 2009. "Case 3487: Megalosaurus crenatissimus Depéret, 1896 (currently Majungasaurus crenatissimus; Dinosauria, Theropoda): proposed replacement of the holotype by a neotype." The Bulletin of Zoological Nomenclature 66 (3):261-264.
  • Carrano, Matthew T. and Sampson, S. D. 2008. "The phylogeny of Ceratosauria (Dinosauria : Theropoda)." Journal of Systematic Palaeontology 6 (2):183-236. https://doi.org/10.1017/S1477201907002246
  • Carrano, Matthew T. 2007. "Walking and running dinosaurs." in Dinosaurs, edited by Holtz, Thomas Richard, Jr., 308. Random House Children's Books.
  • Carrano, Matthew T. 2007. "The appendicular skeleton of Majungasaurus crenatissimus (Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar." Society of Vertebrate Paleontology Memoir 8.Journal of Vertebrate Paleontology 27 (2):163-179.
  • Krause, D. W., Sampson, S. D., Carrano, Matthew T., and O'Conner, Patrick M. 2007. "Overview of the history of discovery, taxonomy, phylogeny, and biogeography of Majungasaurus crenatissimus (Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar." Society of Vertebrate Paleontology Memoir 8.Journal of Vertebrate Paleontology 27 (2):1-20. https://doi.org/10.1671/0272-4634(2007)27[1:OOTHOD]2.0.CO
  • Moreno, Karen, Carrano, Matthew T., and Snyder, Rebecca A. 2007. "Morphological changes in pedal phalanges through ornithopod dinosaur evolution: a biomechanical approach." Journal of Morphology 268 (1):50-63. https://doi.org/10.1002/jmor.10498
  • Carrano, Matthew T. 2006. "Body-size evolution in the Dinosauria." in Amniote Paleobiology: Perspectives on the Evolution of Mammals, Birds, and Reptiles, edited by Carrano, Matthew T., Gaudin, T. J., Blob, R. W., and Wible, J. R., 225-268. University of Chicago Press.
  • Carrano, Matthew T., Gaudin, T. J., Blob, R. W., and Wible, J. R. 2006. Amniote Paleobiology: Perspectives on the Evolution of Mammals, Birds, and Reptiles. University of Chicago Press.
  • Carrano, Matthew T. and Velez-Juarbe, Jorge. 2006. "Paleoecology of the Quarry 9 vertebrate assemblage from Como Bluff Wyoming (Morrison Formation, Late Jurassic)." Palaeogeography palaeoclimatology palaeoecology 237 (2-4):147-159. https://doi.org/10.1016/j.palaeo.2005.11.018
  • Crompton, A. W., Jenkins, F. A., Hopson, S., Gaudin, T. J., and Carrano, Matthew T. 2006. "James Allen Hopson: a biography." in Amniote Paleobiology: Perspectives on the Evolution of Mammals, Birds, and Reptiles, edited by Carrano, Matthew T., Blob, R. W., Gaudin, T., and Wible, J., 507-515. University of Chicago Press.
  • Gaudin, T. J. and Carrano, Matthew T. 2006. "James Allen Hopson: a bibliography (1964-2003)." in Amniote Paleobiology: Perspectives on the Evolution of Mammals, Birds, and Reptiles, edited by Carrano, Matthew T., Blob, R. W., Gaudin, T., and Wible, J., 507-515. University of Chicago Press.
  • Gaudin, T. J., Carrano, Matthew T., Blob, R. W., and Wible, J. R. 2006. "Introduction." in Amniote Paleobiology: Perspectives on the Evolution of Mammals, Birds, and Reptiles, edited by Carrano, Matthew T., Blob, R. W., Gaudin, T., and Wible, J., 1-17. University of Chicago Press.
  • Krause, D. W., O'Connor, P. M., Rasoamiaramanana, A. H., Buckley, G. A., Burney, D., Carrano, Matthew T., Chatrath, P. S., Flynn, J. J., Forster, C. A., Godfrey, L. R., Jungers, W. L., Rogers, R. R., Samonds, K. E., Simons, E. L., and Wyss, A. R. 2006. "Preserving Madagascar's natural heritage: the importance of keeping the island's vertebrate fossils in the public domain." Madagascar Conservation and Development 1 (1):43-47.
  • Carrano, Matthew T. 2005. "The evolution of sauropod locomotion: morphological diversity of a secondarily quadrupedal radiation." in The Sauropods: Evolution and Paleobiology, edited by Curry Rogers, K. A. and Wilson, J. A., 229-251. University of California Press.
  • Carrano, Matthew T., Hutchinson, J. R., and Sampson, S. D. 2005. "New information on Segisaurus halli, a small theropod dinosaur from the Early Jurassic of Arizona." Journal of Vertebrate Paleontology 25 (4):835-849.
  • Carrano, Matthew T. and O'Conner, Patrick M. 2005. "Bird's Eye view." Natural History 114 (4):42-47.
  • O'Keefe, F. Robin and Carrano, Matthew T. 2005. "Correlated trends in the evolution of the plesiosaur locomotor system." Paleobiology 31 (4):656-675.
  • Carrano, Matthew T. 2004. "Why are there no living animals as large as the dinosaurs?." Discover 25 (1):12.
  • Carrano, Matthew T. and Sampson, S. D. 2004. "A review of coelophysoids (Dinosauria: Theropoda) from the Lower Jurassic of Europe, with comments on the late history of the Coelophysoidea." Neues Jahrbuch fAr Geologie und PalAontologie Monatshefte 2004 (9):537-558.
  • Carrano, Matthew T., Sampson, S. D., and Forster, C. A. 2002. "The osteology of Masiakasaurus knopfleri, a small abelisauroid (Dinosauria: Theropoda) from the Late Cretaceous of Madagascar." Journal of Vertebrate Paleontology 22 (3):510-534.

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