Dale Greenwalt

• Ph.D. Iowa State University
• B.A. University of Minnesota, Minneapolis

Visit Dale Greenwalt's Research Site

My work centers on description of the insect fauna of the Coal Creek Member of the Middle Eocene Kishenehn Formation in northwestern Montana. I am also interested in defining the biological and geological processes that resulted in the preservation of these insects and the original biomolecular components that are found in the fossils of the Kishenehn Formation.

Book

• Greenwalt, Dale E. 2023. Remnants of Ancient Life: The New Science of Old Fossils. Princeton, New Jersey: Princeton University Press. https://doi.org/10.2307/j.ctv2sbm8hg.

Journal Article

• Huber, John T. and Greenwalt, Dale E. 2023. "A new compression fossil, Eotriadomeroides abjunctus Huber, gen. & sp. nov. (Hymenoptera, Mymaridae), in Eocene shale from the Kishenehn Formation, USA." Journal of Hymenoptera Research, 96 657–666. https://doi.org/10.3897/jhr.96.107379.
• Smith, MacKenzie A., Greenwalt, Dale E., and Manchester, Steven R. 2023. "Diverse fruits and seeds of the mid-Eocene Kishenehn Formation, northwestern Montana, USA, and their implications for biogeography." Fossil Imprint, 79, (1) 37–88. https://doi.org/10.37520/fi.2023.004.
• Greenwalt, Dale, Haug, Joachim, Harbach, Ralph, and Baranov, Viktor. 2022. "Diversity of culicomorphan dipterans in the Eocene Kishenehn Konservat-Lagerstätte (Montana, USA) and its palaeoecological implications." Palaeontologia Electronica, 25, (1). https://doi.org/10.26879/1165.
• Archibald, S. B., Cannings, Robert A., and Greenwalt, Dale E. 2022. "Kishenehna prima, a new genus and species of darner dragonfly (Odonata, Aeshnidae, Gomphaeschninae) from the early middle Eocene Kishenehn Formation of Montana, USA." Zootaxa, 5099, (4) 496–500. https://doi.org/10.11646/zootaxa.5099.4.5.
• Greenwalt, Dale, Amorim, Dalton, Hauser, Martin, Kerr, Peter, Fitzgerald, Scott, Winterton, Shaun, Cumming, Jeffrey, Evenhuis, Neal, and Sinclair, Bradley. 2022. "Diptera of the Middle Eocene Kishenehn Formation II." Palaeontologia Electronica, https://doi.org/10.26879/1215.
• Amorim, Dalton De Souza and Greenwalt, Dale E. 2020. "Cretaceous and Eocene fossils of the rare extant genus Synneuron Lundstrom (Diptera: Canthyloscledidae): evidence of a true Pangean clade." Cladistics, 36, (4) 413–423. https://doi.org/10.1111/cla.12413.
• Haug, Carolin, Reumer, Jelle W. F., Haug, Joachim T., Arillo, Antonio, Audo, Denis, Azar, Dany, Baranov, Viktor, Beutel, Rolf, Charbonnier, Sylvain, Feldmann, Rodney, Foth, Christian, Fraaije, Rene H. B., Frenzel, Peter, Gasparic, Rok, Greenwalt, Dale E., Harms, Danilo, Hyzny, Matus, Jagt, John W. M., Jagt-Yazykova, Elena A., Jarzembowski, Ed, Kerp, Hans, Kirejtshuk, Alexander G., Klug, Christian, Kopylov, Dmitry S., Kotthoff, Ulrich et al. 2020. "Comment on the letter of the Society of Vertebrate Paleontology (SVP) dated April 21, 2020 regarding "Fossils from conflict zones and reproducibility of fossil-based scientific data": the importance of private collections." Palz, 94 413–429. https://doi.org/10.1007/s12542-020-00522-x.
• Greenwalt, Dale E. and Blagoderov, Vladimir A. 2019. "Review of the fossil record of Bolitophilidae, with description of new taxa and discussion of position of Mangas Kovalev (Diptera: Sciaroidea)." Zootaxa, 4567, (3) 546–560. https://doi.org/10.11646/zootaxa.4567.3.6.
• Greenwalt, Dale, Bickel, Daniel, Kerr, Peter, Curler, Gregory, Brown, Brian, de Jong, Herman, Fitzgerald, Scott, Dikow, Torsten, Tkoč, Michal, Kehlmaier, Christian, and Amorim, Dalton De Souza. 2019. "Diptera of the middle Eocene Kishenehn Formation. I. Documentation of diversity at the family level." Palaeontologia Electronica, 22.2.50. https://doi.org/10.26879/891.
• Greenwalt, Dale E. and Moulton, John K. 2016. "The first fossil New World Dixidae with a critical discussion of generic definitions." Palaeontologia Electronica, 19, (3) 55A–55A. https://doi.org/10.26879/656.
• Greenwalt, Dale, Rose, Timothy R., and Chatzimanolis, Stylianos. 2016. "Preservation of mandibular zinc in a beetle from the Eocene Kishenehn Formation of Montana, USA." Canadian Journal of Earth Sciences, 1–8. https://doi.org/10.1139/cjes-2015-0157.
• Greenwalt, Dale E., Rose, Tim R., Siljestrom, Sandra M., Goreva, Yulia S., Constenius, Kurt N., and Wingerath, Jonathan G. 2015. "Taphonomy of the fossil insects of the Middle Eocene Kishenehn Formation." Acta Palaeontologica Polonica, 60, (4) 931–947. https://doi.org/10.4202/app.00071.2014.
• Greenwalt, Dale E. and Vidlička, Lubomír. 2015. "Latiblattella avita sp. nov. (Blattaria: Ectobiidae) from the Eocene Kishenehn Formation, Montana, USA." Palaeontologia Electronica, 18.1.16A. https://doi.org/10.26879/511.
• Greenwalt, Dale E., Wingerath, Jonathan G., and Evenhuis, Neal L. 2015. "Two new and disparate fossil bee flies (Bombyliidae: Anthracinae) from the Americas and reassessment of Anthrax dentoni Lewis, 1969." Palaeontologia Electronica, 18, (3) 1–10. https://doi.org/10.26879/582.
• LaPolla, John S. and Greenwalt, Dale E. 2015. "Fossil Ants (Hymenoptera: Formicidae) of the Middle Eocene Kishenehn Formation." Sociobiology, 62, (2) 163–174. https://doi.org/10.13102/sociobiology.v62i2.163-174.
• Greenwalt, Dale E. and Bechly, Günter. 2014. "A re-description of the fossil damselfly Eolestes syntheticus Cockerell, 1940 (Odonata: Zygoptera: Eolestidae n. fam.) with description of new taxa from the Eocene of North America." Zootaxa, 3887, (2) 138–156. https://doi.org/10.11646/zootaxa.3887.2.2.
• Greenwalt, Dale E. and Engel, Michael S. 2014. "A diminutive pelecinid wasp from the Eocene Kishenehn Formation of northwestern Montana (Hymenoptera: Pelecinidae)." Novitates Paleoentomologicae, 8 1–9.
• Greenwalt, Dale E. and Rust, Jes. 2014. "A new species of Pseudotettigonia Zeuner (Orthoptera: Tettigoniidae) with an intact stridulatory field and reexamination of the subfamily Pseudotettigoniinae." Systematic Entomology, 39, (2) 256–263. https://doi.org/10.1111/syen.12050.
• Greenwalt, Dale E., Goreva, Yulia S., Siljeström, Sandra M., Rose, Tim, and Harbach, Ralph E. 2013. "Hemoglobin-derived porphyrins preserved in a Middle Eocene blood-engorged mosquito." Proceedings of the National Academy of Sciences of the United States of America, 110, (46) 18496–18500. https://doi.org/10.1073/pnas.1310885110.
• Greenwalt, Dale E. and Labandeira, Conrad C. 2013. "The Amazing Fossil Insects of the Eocene Kishenehn Formation in NW Montana." Rocks & Minerals, 88, (5) 434–441. https://doi.org/10.1080/00357529.2013.809972.
• Ouvrard, D., Burckhardt, D., and Greenwalt, Dale E. 2013. "The oldest jumping plant-louse (Insecta: Hemiptera: Sternorrhyncha) with comments on the classification and nomenclature of the Palaeogene Psylloidea." Acta Musei Moraviae, 98, (2) 21–33.
• Shockley, Floyd W. and Greenwalt, Dale E. 2013. "Ptenidium kishenehnicum (Coleoptera: Ptiliidae), a new fossil described from the Kishenehn oil shales, with a checklist of previously known fossil ptiliids." Proceedings of the Entomological Society of Washington, 115, (2) 173–181. https://doi.org/10.4289/0013-8797.115.2.173.
• Greenwalt, D. and Labandeira, Conrad C. 2012. "Eocene Fossil Insects from the Kishenehn Formation, Flathead National Forest." Diggin' Deep, 300, (3-4).
• Harbach, Ralph E. and Greenwalt, Dale E. 2012. "Two Eocene species of Culiseta (Diptera: Culicidae) from the Kishenehn Formation in Montana." Zootaxa, (3530) 25–34.
• Huber, John T. and Greenwalt, Dale E. 2011. "Compression fossil Mymaridae (Hymenoptera) from Kishenehn oil shales, with description of two new genera and review of Tertiary amber genera." Zookeys, (130) 473–494. https://doi.org/10.3897/zookeys.130.1717.