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Smithsonian National Museum of Natural History

Wayne Mathis

Research Entomologist, Diptera (retired)
Department / Division
Entomology
Education
  • BS Brigham Young 
  • PhD Oregon State
Research Interests

Systematics, biology, and zoogeography of Canacidae, and Ephydroidea, with special emphasis on Ephydridae

For over 40 years, I have focused on the systematics and evolution of higher Diptera (true flies), especially families in the Ephydroidea and Opomyzoidea. My research primarily employs the methodology of cladistic analysis to unravel, reconstruct, and document phylogenetic relationships and how these relationships impinge on historical biogeography. The phylogenetic investigations and analyses run the gamut from species-level revisions to basal lineages of superfamilies. Documentation includes comprehensive systematics studies at all levels of the classification, faunistic studies, catalogs and conspectuses. Although my comprehensive studies are global, I have focused field work in the neotropics, especially the Caribbean, and on islands, such as Sri Lanka, the Seychelles, New Zealand, and the barrier islands of Belize.

for more information: Research Gate

Publications
Journal Article
  • Murphy, William L., Mathis, Wayne N., and Knutson, Lloyd V. 2024. "Revision of Clam-Killing Flies of the Genus Renocera Hendel (Diptera: Sciomyzidae), with Emphasis on Species in the Nearctic Region." Proceedings of the Entomological Society of Washington, 125, (3). https://doi.org/10.4289/0013-8797.125.3.326.
  • Costa, Sávio Cunha, Pirani, Gabriela, and Mathis, Wayne N. 2023. "First record of Diastatidae (Diptera: Ephydroidea) from Brazil, with the description of a new species." Zootaxa, 5278, (2) 378–386. https://doi.org/10.11646/zootaxa.5278.2.10.
  • Mathis, Wayne N. and Zatwarnicki, Tadeusz. 2023. "Phylogeny and Taxonomy of the Risa Genus-group (Diptera: Ephydridae), with Description of a New Genus from Australia." Records of the Australian Museum, 75, (1) 1–23. https://doi.org/10.3853/j.2201-4349.75.2023.1823.
  • Mathis, Wayne N. and Zatwarnicki, Tadeusz. 2022. "Biogeographic Anomalies in Shore Flies as Revealed in Revision of the Shore-Fly Genus Subpelignus Papp (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 124, (3). https://doi.org/10.4289/0013-8797.124.3.683.
  • Murphy, William L. and Mathis, Wayne N. 2021. "Bibliography for Comprehensive taxonomic, faunistic, biological, and geographic inventory and analysis of the Sciomyzidae (Diptera: Acalyptratae) of the Delmarva region and nearby states in eastern North America." Zootaxa, 5016, (2) 289–293. https://doi.org/10.11646/zootaxa.5016.2.9.
  • Costa, Daniel N. R., Mathis, Wayne N., and Marinoni, Luciane. 2019. "Revision of Scatella (Neoscatella) (Diptera: Ephydridae) from the New Zealand subregion." Austral Entomology, 58, (1) 85–95. https://doi.org/10.1111/aen.12316.
  • Mathis, Wayne N. and Zatwarnicki, Tadeusz. 2019. "Revision of the Shore-fly Genera Parahyadina Tonnoir and Malloch and New Zealand Hyadina Haliday (Diptera: Ephydridae)." Zootaxa, 4623, (3) 401–440. https://doi.org/10.11646/zootaxa.4623.3.1.
  • Mathis, Wayne N. and Zatwarnicki, Tadeusz. 2019. "New taxonomic twists in shore flies as revealed in discovery of Neozealides, a new genus from New Zealand (Diptera: Ephydridae)." Zootaxa, 4656, (3) 487–500. https://doi.org/10.11646/zootaxa.4656.3.6.
  • Zatwarnicki, Tadeusz and Mathis, Wayne N. 2019. "New discoveries in New World Atissini as revealed in revision of Pelignellus Sturtevant & Wheeler (Diptera: Ephydridae)." Israel Journal of Entomology, 49, (2) 11–26. https://doi.org/10.5281/zenodo.3244845.
  • Borkent, Art, Brown, Brian V., Adler, Peter H., Amorim, Dalton De Souza, Barber, Kevin, Bickel, Daniel, Boucher, Stephanie, Brooks, Scott E., Burger, John, Burington, Z. L., Capellari, Renato S., Costa, Daniel N. R., Cumming, Jeffrey M., Curler, Greg, Dick, Carl W., Epler, J. H., Fisher, Eric, Gaimari, Stephen D., Gelhaus, Jon, Grimaldi, David A., Hash, John, Hauser, Martin, Hippa, Heikki, Ibanez-Bernal, Sergio, Jaschhof, Mathias et al. 2018. "Remarkable fly (Diptera) diversity in a patch of Costa Rican cloud forest: Why inventory is a vital science." Zootaxa, 4402, (1) 53–90. https://doi.org/10.11646/zootaxa.4402.1.3.
  • Brown, Brian V., Borkent, Art, Adler, Peter H., Amorim, Dalton De Souza, Barber, Kevin, Bickel, Daniel, Boucher, Stephanie, Brooks, Scott E., Burger, John, Burington, Zelia L., Capellari, Renato S., Costa, Daniel N. R., Cumming, Jeffrey M., Curler, Greg, Dick, Carl W., Epler, John H., Fisher, Eric, Gaimari, Stephen D., Gelhaus, Jon, Grimaldi, David A., Hash, John, Hauser, Martin, Hippa, Heikki, Ibáñez-Bernal, Sergio, Jaschhof, Mathias et al. 2018. "Comprehensive inventory of true flies (Diptera) at a tropical site." Communications Biology, 1, (1). https://doi.org/10.1038/s42003-018-0022-x.
  • Murphy, William L., Mathis, Wayne Neilsen, and Knutson, Lloyd, V. 2018. "Comprehensive taxonomic, faunistic, biological, and geographic inventory and analysis of the Sciomyzidae (Diptera: Acalyptratae) of the Delmarva region and nearby states in eastern North America." Zootaxa, 4430, (1) 1–299. https://doi.org/10.11646/zootaxa.4430.1.1.
  • Costa, Daniel N. R., Mathis, Wayne N., and Marinoni, Luciane. 2016. "A revision of the shore-fly genus Lamproclasiopa Hendel (Diptera, Ephydridae)." ZooKeys, (631) 1–99. https://doi.org/10.3897/zookeys.631.10718.
  • Costa, Daniel N. R., Savaris, Marcoandre, Marinoni, Luciane, and Mathis, Wayne N. 2016. "Two new, brachypterous Limnellia species from the Venezuelan Andes (Diptera: Ephydridae)." Zootaxa, 4144, (3) 301–315. https://doi.org/10.11646/zootaxa.4144.3.1.
  • Mathis, Wayne N., Costa, Daniel N. R., and Marinoni, Luciane. 2016. "A review of the Neotropical species of the shore-fly genera Orasiopa and Pectinifer (Diptera: Ephydridae)." Sociedade Brasileira de Zoologia, 33, (3). https://doi.org/10.1590/S1984-4689zool-20150169.
  • Mathis, Wayne N. and Marinoni, Luciane. 2016. "Revision of Ephydrini Zetterstedt (Diptera: Ephydridae) from the Americas south of the United States." Zootaxa, 4116, (1) 1–110. https://doi.org/10.11646/zootaxa.4116.1.
  • Mathis, Wayne Neilsen, Júnior, Francisco De Assis Rodrigues, and Couri, Márcia S. 2016. "Family Ephydridae." Zootaxa, 4122, (1) 752–770. https://doi.org/10.11646/zootaxa.4122.1.64.
  • Savaris, Marcoandre, Marinoni, Luciane, and Mathis, Wayne Neilsen. 2016. "Revision of the shore-fly genus Peltopsilopa Hendel (Diptera: Ephydridae)." Zootaxa, 4083, (1) 83–98. https://doi.org/10.11646/zootaxa.4083.1.4.
  • Assis Rodrigues Junior, Francisco De, Mathis, Wayne Neilsen, and Hauser, Martin. 2015. "Argentine Hydrellia Robineau-Desvoidy (Diptera, Ephydridae): new species and key to identification." Zootaxa, 3957, (1) 131–136. https://doi.org/10.11646/zootaxa.3957.1.11.
  • Costa, Daniel N. R., Mathis, Wayne Neilsen, and Marinoni, Luciane. 2015. "Morphological phylogenetic analysis confirms the monophyly of Rhysophora Cresson (Diptera: Ephydridae) and provides a framework for a revision of the genus." Invertebrate Systematics, 29, (1) 71–103. https://doi.org/10.1071/IS14028.
  • Mathis, Wayne Neilsen, Costa, Daniel N. R., and Marinoni, Luciane. 2015. "A review of Mimapsilopa Cresson (Diptera: Ephydridae) from Brazil." Zootaxa, 3926, (4) 499–522. https://doi.org/10.11646/zootaxa.3926.4.3.
  • Pape, Thomas, Beuk, Paul, Pont, Adrian, Shatalkin, Anatole, Ozerov, Andrey, Woznica, Andrzej, Merz, Bernhard, Bystrowski, Cezary, Raper, Chris, Bergström, Christer, Kehlmaier, Christian, Clements, David, Greathead, David, Kameneva, Elena, Nartshuk, Emilia, Petersen, Frederik, Weber, Gisela, Bächli, Gerhard, Geller-Grimm, Fritz, Van de Weyer, Guy, Tschorsnig, Hans-Peter, de Jong, Herman, van Zuijlen, Jan-Willem, Vanhara, Jaromír, Rohácek, Jindrich et al. 2015. "Fauna Europaea: Diptera - Brachycera." Biodiversity Data Journal, 3. https://doi.org/10.3897/BDJ.3.e4187.
  • Ale-Rocha, Rosaly, Freitas, Geovania, and Mathis, Wayne Neilsen. 2014. "Revision of the Neotropical genus Marbenia Malloch (Diptera: Periscelididae)." Zootaxa, 3872, (4) 365–375. https://doi.org/10.11646/zootaxa.3872.4.3.
  • Mathis, Wayne Neilsen, Marinoni, Luciane, and Costa, Daniel N. R. 2014. "A review of Scatellini (Diptera: Ephydridae) from Brazil." Zoologia (Curitiba), 31, (6) 561–576. https://doi.org/10.1590/S1984-46702014000600005.
  • Rodrigues Junior, Francisco De Assis, Mathis, Wayne Neilsen, and Couri, Marcia Souto. 2014. "Hydrellia Robineau-Desvoidy (Diptera: Ephydridae) from Brazil with an emphasis on the faunas from the states of Parana and Rio de Janeiro." Zootaxa, 3753, (6) 501–541. https://doi.org/10.11646/zootaxa.3753.6.1.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 2013. "A revision of the shore-fly genus Hydrochasma Hendel (Diptera, Ephydridae)." ZooKeys, 363 1–161. https://doi.org/10.3897/zookeys.363.6482.
  • Huang, Yiau-Min, Mathis, Wayne Neilsen, and Wilkerson, Richard C. 2012. "Subgeneric Position of Aedes dufouri Hamon with Notes on the Subgenus Levua Stone and Bohart (Diptera: Culicidae)." Proceedings of the Entomological Society of Washington, 114, (2) 194–204. https://doi.org/10.4289/0013-8797.114.2.194.
  • Mathis, Wayne Neilsen and Freidberg, Amnon. 2012. "Periscelis stuckenbergi sp. n., the first record of the genus from the Afrotropical Region (Diptera: Periscelididae: Periscelidinae)." African Invertebrates, 53, (1) 231–238. https://doi.org/10.5733/afin.053.0115.
  • Mathis, Wayne Neilsen and Marinoni, Luciane. 2012. "A conspectus on the Canacidae (Diptera) of Brazil." ZooKeys, 162 59–92. https://doi.org/10.3897/zookeys.162.2370.
  • Mathis, Wayne Neilsen, Rung, Alessandra, and Kotrba, Marion. 2012. "A revision of the genus Planinasus Cresson (Diptera, Periscelididae)." ZooKeys, 225 1–83. https://doi.org/10.3897/zookeys.225.3721.
  • Mathis, Wayne Neilsen, Steinly, Bruce A., Jr., Buckingham, Gary R., and Deonier, Bryan C. 2012. "Dick L. Deonier 1936-2011 Obituary." Proceedings of the Entomological Society of Washington, 114, (1) 169–172. https://doi.org/10.4289/082.114.0103.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 2012. "A revision of the new world species of Gymnoclasiopa Hendel (Diptera, Ephydridae)." ZooKeys, 248 1–69. https://doi.org/10.3897/zookeys.248.4106.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 2012. "Review of the shore-fly genus Oedenopiforma (Diptera: Ephydridae)." The Canadian Entomologist, 144, (1) 81–92. https://doi.org/10.4039/tce.2012.9.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 2012. "A revision of the new world species of Polytrichophora Cresson and Facitrichophora, new genus (Diptera, Ephydridae)." ZooKeys, 231 1–116. https://doi.org/10.3897/zookeys.231.3687.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 2012. "Revision of New World Species of the Shore-fly Subgenus Allotrichoma Becker of the Genus Allotrichoma with Description of the Subgenus Neotrichoma (Diptera, Ephydridae, Hecamedini)." ZooKeys, 161 1–101. https://doi.org/10.3897/zookeys.161.2016.
  • Zhang, Junhua, Yang, Ding, and Mathis, Wayne Neilsen. 2012. "A review of the species of Rhynchopsilopa Hendel from China (Diptera, Ephydridae)." Zookeys, (216) 23–42. https://doi.org/10.3897/zookeys.216.3224.
  • Gaimari, Stephen D. and Mathis, Wayne Neilsen. 2011. "World catalog and conspectus on the family Odiniidae (Diptera: Schizophora)." Myia, 12 291–339.
  • Mathis, Wayne Neilsen. 2011. "World catalog and conspectus on the family Heterocheilidae (Diptera: Schizophora)." Myia, 12 281–289.
  • Mathis, Wayne Neilsen. 2011. "World Catalog and Conspectus on the Family Helcomyzidae (Diptera: Schizophora)." Myia, 12 267–280.
  • Mathis, Wayne Neilsen and Barraclough, D. A. 2011. "World catalog and conspectus on the family Diastatidae (Diptera: Schizophora)." Myia, 12 235–266.
  • Mathis, Wayne Neilsen and Marinoni, Luciane. 2011. "A review of Parydra (Diptera: Ephydridae) from Brazil." Zoologia, 28, (4) 505–512. https://doi.org/10.1590/S1984-46702011000400012.
  • Mathis, Wayne Neilsen and McAlpine, D. K. 2011. "A catalog and conspectus on the family Coelopidae (Diptera: Schizophora)." Myia, 12 171–205.
  • Mathis, Wayne Neilsen and Rung, A. 2011. "World catalog and conspectus on the family Periscelididae (Diptera: Schizophora)." Myia, 12 341–377.
  • Mathis, Wayne Neilsen and Sueyoshi, M. 2011. "World catalog and conspectus on the family Dryomyzidae (Diptera: Schizophora)." Myia, 12 207–233.
  • Mathis, Wayne Neilsen and Sueyoshi, Masahiro. 2011. "New species of the genus Cyamops Melander from New Zealand (Diptera, Periscelididae, Stenomicrinae)." ZooKeys, 114 28–40. https://doi.org/10.3897/zookeys.114.1310.
  • Huang, Yiau-Min, Mathis, Wayne Neilsen, and Wilkerson, Richard C. 2010. "Coetzeemyia, a new subgenus of Aedes, and a redescription of the holotype female of Aedes (Coetzeemyia) fryeri (Theobald) (Diptera: Culicidae)." Zootaxa, (2638) 1–24.
  • Krivosheina, Marina and Mathis, Wayne Neilsen. 2010. "A Review of the Shore-Fly Genus Axysta Haliday (Diptera: Ephydridae) with Description of New Species from Australia, India, and Thailand." Entomological News, 121, (4) 369–390. https://doi.org/10.3157/021.121.0412.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 2010. "New Species and Taxonomic Clarifications for Shore Flies from the Delmarva States (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 112, (1) 97–128. https://doi.org/10.4289/0013-8797.112.1.97.
  • Munari, Lorenzo and Mathis, Wayne Neilsen. 2010. "World Catalog of the Family Canacidae (including Tethinidae) (Diptera), with keys to the supraspecific taxa." Zootaxa, (2471) 1–84.
  • Myers, Janet and Mathis, Wayne Neilsen. 2010. "Hollis (Holly) Barton Williams 1951-2009." Proceedings of the Entomological Society of Washington, 112, (3) 481–482. https://doi.org/10.4289/0013-8797-112.3.481.
  • Zatwarnicki, Tadeusz and Mathis, Wayne Neilsen. 2010. "A Revision of the Nivea Group of the Shore-fly Genus Ditrichophora Cresson (Diptera: Ephydridae)." Transactions of the American Entomological Society, 136, (3-4) 199–215. https://doi.org/10.3157/061.136.0301.
  • Kotrba, Marion and Mathis, Wayne Neilsen. 2009. "The Internal Female Reproductive Tract of the Enigmatic Genus Risa (Diptera: Schizophora: Ephydroidea) and its Phylogenetic Implications." Proceedings of the Entomological Society of Washington, 111, (3) 627–640. https://doi.org/10.4289/0013-8797-111.3.627.
  • Mathis, Wayne Neilsen, Knutson, Lloyd V., and Murphy, William L. 2009. "A New Species of Snail-Killing Fly of the Genus Dictya Meigen from the Delmarva States (Diptera: Sciomyzidae)." Proceedings of the Entomological Society of Washington, 111, (4) 785–794. https://doi.org/10.4289/0013-8797-111.4.785.
  • Mathis, Wayne Neilsen and Mathis, Dianne. 2009. "Species of the Shore-Fly Genus Glenanthe Haliday (Diptera: Ephydridae) from New Mexico." Proceedings of the Entomological Society of Washington, 111, (2) 454–463. https://doi.org/10.4289/0013-8797-111.2.454.
  • Mathis, Wayne Neilsen, Zatwarnicki, Tadeusz, and Kubatova-Hirsova, Hana. 2009. "A revision of the shore-fly genus Philotelma Becker (Diptera: Ephydridae)." Insect Systematics & Evolution, 40, (2) 121–158. https://doi.org/10.1163/187631209X416714.
  • Zhang, Junhua, Yang, Ding, and Mathis, Wayne Neilsen. 2009. "A New Species of the Shore-Fly Genus Oedenopiforma Cogan from the Oriental Region, with an Updated Key to the Species (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 111, (1) 199–203. https://doi.org/10.4289/0013-8797-111.1.199.
  • Foster, G. A. and Mathis, Wayne Neilsen. 2008. "Review of the genus Tethina Haliday (Diptera : Canacidae : Tethininae) from western North America." Proceedings of the Entomological Society of Washington, 110, (2) 300–330. https://doi.org/10.4289/07-059.1.
  • Foster, G. A. and Mathis, Wayne Neilsen. 2008. "A review of the Tethininae (Diptera : Canacidae) from the Galapagos Islands." Proceedings of the Entomological Society of Washington, 110, (3) 743–752. https://doi.org/10.4289/07-078.1.
  • Mathis, Wayne Neilsen. 2008. "Two new Neotropical genera of the shore-fly tribe Ephydrini Zetterstedt (Diptera: Ephydridae)." Zootaxa, 1874 1–15.
  • Mathis, Wayne Neilsen and Ohishi, H. 2008. "Revision of the shore-fly genus Eremomusca mathis (Diptera : Ephydridae) with description of a new species from Japan." Proceedings of the Entomological Society of Washington, 110, (1) 67–78. https://doi.org/10.4289/0013-8797-110.1.67.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 2008. "A Revision of the Nearctic Species of Hecamedoides Hendel (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 110, (4) 1012–1027. https://doi.org/10.4289/0013-8797-110.4.1012.
  • Brake, Irina and Mathis, Wayne Neilsen. 2007. "Revision of the genus Australimyza Harrison (Diptera: Australimyzidae)." Systematic Entomology, 32, (2) 252–275. https://doi.org/10.1111/j.1365-3113.2006.00363.x.
  • Edmiston, J. F. and Mathis, Wayne Neilsen. 2007. "New Zealand species of the shore-fly genus Nostima Coquillett (Diptera : Ephydridae)." Zootaxa, (1661) 1–16.
  • Mathis, Wayne Neilsen and Foster, G. A. 2007. "Canacidae (Diptera) from the Delmarva States." Proceedings of the Biological Society of Washington, 120, (4) 387–428. https://doi.org/10.2988/0006-324X(2007)120[387:CDFTDS]2.0.CO;2.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 2007. "A revisiion of the species of the genus Dichaeta Meigen (Diptera: Ephydridae)." Annales Zoologici, 57, (4) 783–822.
  • Zatwarnicki, Tadeusz and Mathis, Wayne Neilsen. 2007. "A revision of the Palearctic species of the shore-fly genus Discomyza Meigen (Diptera : Ephydridae)." Insect Systematics & Evolution, 38, (3) 241–266. https://doi.org/10.1163/187631207788754448.
  • Zhang, J. J., Yang, D., and Mathis, Wayne Neilsen. 2007. "Species of the genus Zeros Cresson (Diptera : Ephydridae) from China." Proceedings of the Entomological Society of Washington, 109, (4) 872–879.
  • Marinoni, Luciane and Mathis, Wayne Neilsen. 2006. "A cladistic analysis of the Neotropoical genus Sepedonea Steyskal (Diptera: Sciomyzidae)." Zootaxa, 1236 37–52.
  • Mathis, Wayne Neilsen, Castro, B. A., Wasy, M. O., and Zatwarnicki, Tadeusz. 2006. "Discovery of the South American rice miner (Diptera: Ephydridae) in the Southeastern United States." Annals of the Entomological Society of America, 99, (6) 999–1005. https://doi.org/10.1603/0013-8746(2006)99[999:DOTSAR]2.0.CO;2.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 2006. "A review of the New World species of the shore-fly genus Leptopsilopa Cresson (Diptera: Ephydridae)." Annales Zoologica, 56, (1) 85–138.
  • Edmiston, J. F. and Mathis, Wayne Neilsen. 2005. "A review of two Nearctic species of the shore-fly genus Philygria Stenhammar, P. debilis Loew and P. nigrescens (Cresson) (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 107, (1) 7–20.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 2005. "Revision of New World species of shore fly genus Discomyza Meigen (Diptera: Ephydridae." Annals of the Entomological Society of America, 98, (4) 431–443.
  • Mathis, Wayne Neilsen and Zumbado, M. A. 2005. "Description of Scatella savegre, a new species from Costa Rica in the triseta group (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 107, (2) 386–391.
  • Rung, A., Mathis, Wayne Neilsen, and Papp, L. 2005. "Curiosimusca, gen. nov., and three new species in the family Aulacigastridae from the Oriental Region (Diptera: Opomyzoidea)." Zootaxa, 1009 21–36.
  • Zhang, J., Yang, D., and Mathis, Wayne Neilsen. 2005. "A new genus and species of Ephydridae (Diptera) from the Oriental Region." Zootaxa, 1040 31–43.
  • Mathis, Wayne Neilsen and Rung, Alessandra. 2004. "Redescription of Diopsosoma primum Malloch (Diptera: Periscelididae)." Revista Brasileira de Entomologia, 48, (3) 303–309.
  • Mathis, Wayne Neilsen and Tadeusz, Zatwarnicki. 2004. "A review of the species of Hyadina Haliday (Diptera: Ephydridae) occurring in Israel." Israel Journal of Entomology, 34 35–57.
  • Mathis, Wayne Neilsen and Tadeusz, Zatwarnicki. 2004. "Tronamyia, a new genus of shore-flies and a phylogenetic reassessment of the tribe Lipochaetini Becker (Diptera: Ephydridae)." Annals of the Entomological Society of America, 97, (2) 259–270.
  • Mathis, Wayne Neilsen and Tadeusz, Zatwarnicki. 2004. "A revision of the shore-fly genus Cressonomyia Arnaud (Diptera: Ephydridae), with comments on species that have been excluded." Proceedings of the Entomological Society of Washington, 106, (2) 249–279.
  • Mathis, Wayne Neilsen, Tadeusz, Zatwarnicki, and John, W. M. M. 2004. "A review of unreported shore-fly genera of the tribe Scatellini from the New Zealand subregion (Diptera: Ephydridae)." Zootaxa, (622) 1–27.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 2004. "A revision of the shore-fly genus Trimerogastra Hendel (Diptera: Ephydridae). Pages 89-108. In N. L. Evenhuis and K. Y. Kaneshiro, eds., D. Elmo Hardy Memorial Volume. Contributions to the Systematics and Evolution of Diptera." Bishop Museum Bulletins in Entomology, 12 89–108.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 2004. "A review of two Nearctic shore-fly species in the genus Psilopa Fallén that were included in the genus Cressonomyia Arnaud (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 106, (3) 639–648.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 2004. "Description of a new species of Hyadina Haliday from southern California (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 106, (3) 705–716.
  • Steyskal, George C., Thompson, F. Christian, Mathis, Wayne Neilsen, and Knutson, Lloyd. 2004. "The type species of Ilione (Diptera: Sciomyzidae)." Studia Dipterologica, 10, (2) 559–564.
  • Sueyoshi, Masahiro and Mathis, Wayne Neilsen. 2004. "A new species of Cyamops Melander, 1913 (Diptera: Periscelididae) from Japan and a review of the Japanese Periscelididae." Proceedings of the Entomological Society of Washington, 106, (1) 74–84.
  • Cheng, Lanna and Mathis, Wayne Neilsen. 2003. "Marine insects of Guam: Heteroptera and Diptera." Micronesica, 35-36 514–522.
  • Mathis, Wayne Neilsen and Michelle, D. Trautwein. 2003. "A revision and phylogenetic study of Lipochaeta Coquillett (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 105, (1) 30–49.
  • Mathis, Wayne Neilsen and Winkler, Isaac S. 2003. "A review of the Neotropical species of Brachydeutera Loew (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 105, (2) 407–424.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 2003. "Shore flies of the Republic of Seychelles (Diptera: Ephydridae)." Annales Zoologici (Warszawa), 53, (4) 585–650.
  • Dikow, Torsten and Mathis, Wayne Neilsen. 2002. "A revision and phylogenetic study of Actocetor Becker (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 104, (2) 249–290.
  • Freidberg, Amnon and Mathis, Wayne Neilsen. 2002. "The Periscelididae (Diptera) of Israel, with description of a new species of Stenomicra Coquillett." Israel journal of zoology, 48 43–51.
  • Kirk-Spriggs, A. H., Ismay, J. W., Ackland, M., Rohácek, J., Mathis, Wayne Neilsen, Foster, G. A., Pape, T., Cranston, P. S., and Meier, Rudolf. 2001. "Inter-tidal Diptera of southwestern Africa (Chironomidae, Canacidae, Chloropidae, Milichiidae, Tethinidae, Ephydridae, Sphaeroceridae, Coelopidae, Sarcophagidae and Anthomyiidae)." Cimbebasia, 17 85–135.
  • Papp, L. and Mathis, Wayne Neilsen. 2001. "A review of the family Nannodastiidae (Diptera)." Proceedings of the Entomological Society of Washington, 103, (2) 337–348.
  • Zatwarnicki, Tadeusz and Mathis, Wayne Neilsen. 2001. "A generic classification of the tribe Discocerinini (Diptera: Ephydridae)." Annales Zoologici, 51, (1) 1–51.
  • Baptista, Alessandra R. P. and Mathis, Wayne Neilsen. 2000. "Notes on the genus Cyamops Melander (Diptera: Periscelididae), including description of ten new species." Proceedings of the Entomological Society of Washington, 102, (3) 481–506.
  • Foster, George A. and Mathis, Wayne Neilsen. 2000. "Notes on the Neotropical species of Tethina Haliday (Diptera: Tethinidae)." Proceedings of the Entomological Society of Washington, 102, (33) 542–548.
  • Marinoni, Luciane and Mathis, Wayne Neilsen. 2000. "A cladistic analysis of Sciomyzidae Fallén (Diptera)." Proceedings of the Biological Society of Washington, 113, (1) 162–209.
  • Mathis, Wayne Neilsen and Edmiston, J. F. 2000. "A revision of the shore-fly genus Lemnaphila Cresson (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 102 652–677.
  • Mathis, Wayne Neilsen. 1999. "A Review of the Beach-Fly Genus Isocance Mathis (Diptera: Canacidae)." Proceedings of the Entomological Society of Washington, 101, (2) 347–358.
  • Foster, George A. and Mathis, Wayne Neilsen. 1998. "A Revision of the Family Tethinidae (Diptera) from the Caribbean, Gulf of Mexico, and Bermuda." Proceedings of the Entomological Society of Washington, 100, (4) 601–632.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 1998. "A Review of the West Indian Species of Mimapsilopa Cresson (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 100, (1) 7–24.
  • Feller, Ilka C. and Mathis, Wayne Neilsen. 1997. "Primary Herbivory by Wood-Boring Insects along an Architectural Gradient of Rhizophora mangle." Biotropica, 29, (4) 440–451.
  • Mathis, Wayne Neilsen. 1997. "A Review of the Shore-Fly Genus Diphuia Cresson (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 99, (1) 28–36.
  • Mathis, Wayne Neilsen. 1997. "A Review of the Hoguei Group of Paracanace Mathis and Wirth (Diptera: Canacidae)." Memoirs of the Entomological Society of Washington, 18 140–148.
  • Mathis, Wayne Neilsen. 1997. "A Revision of Neotropical Ditricohphora Cresson (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 99 -num-4-num- 697–704.
  • Baptista, A. R. P. and Mathis, Wayne Neilsen. 1996. "A New Species of Cyamops Melander (Diptera: Periscelididae) from Brazil, with Distributional Notes on Another Species." Proceedings of the Entomological Society of Washington, 98, (2) 245–248.
  • Mathis, Wayne Neilsen. 1996. "Australian beach flies (Diptera: Canacidae)." Proceedings of the Biological Society of Washington, 109, (2) 326–348.
  • Mathis, Wayne Neilsen. 1995. "Shore Flies of the GalApagos Islands (Diptera: Ephydridae)." Annals of the Entomological Society of America, 88, (5) 627–640.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 1995. "World Catalog of Shore Flies (Diptera: Ephydridae)." Memoirs on Entomology, International, Associated Publishers, 4 1–423.
  • Rao, N. and Mathis, Wayne Neilsen. 1995. "A Revision of the Shore-fly Genus Rhysophora Cresson (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 97, (3) 666–688.
  • Mathis, Wayne Neilsen. 1994. "A Revision of the Shore-Fly Genus Clasiopella Hendel (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 96, (3) 454–465.
  • Mathis, Wayne Neilsen and Freidberg, Amnon. 1994. "A Revision of the Neartic Aulacigaster MacQuart with Notes on A. Leucopeze (Meigen) from the Paleartic Region (Diptera: Aulacigastridae)." Proceedings of the Entomological Society of Washington, 96, (4) 583–598.
  • Zatwarnicki, Tadeusz and Mathis, Wayne Neilsen. 1994. "Phylogeny and Classification of the Genus Scatophila Becker (Diptera: Ephydirdae)." Annales de la Societe Entomologique de France, 29, (4) 351–370.
  • Evenhuis, Neal L., Mathis, Wayne Neilsen, and Thompson, F. Christian. 1993. "Comment on the Proposal to Conserve Oecothea Haliday in Curtis, 1836 and to Designate Helomyza fenestralis Fallen, 1820 as the Type Species (Insect, Diptera)." The Bulletin of Zoological Nomenclature, 50, (3) 235–236.
  • Grimaldi, D. A. and Mathis, Wayne Neilsen. 1993. "Fossil Periscelididae (Diptera)." Proceedings of the Entomological Society of Washington, 95, (3) 383–403.
  • Mathis, Wayne Neilsen. 1993. "A New Species and Subgenus of Periscelis Loew from Australia (Diptera: Periscelididae)." Journal of the Australian Entomological Society, 32 13–19.
  • Mathis, Wayne Neilsen. 1993. "A Revision of the Shore-Fly Genera Hostis Cresson and Paratissa Coquillett (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 95, (1) 21–47.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 1993. "A Revision of the Shore-Fly Genus Chlorichaeta Becker (Diptera: Ephydridae)." Tijdschrift voor Entomologie, 136, (1) 55–76.
  • Mathis, Wayne Neilsen. 1992. "The First Shore Fly of the Genus Glenanthe Haliday from the Australasian Region (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 94, (1) 78–82.
  • Mathis, Wayne Neilsen and Papp, L. 1992. "A New Genus of Periscelididae (Diptera) from the Neotropics." Proceedings of the Biological Society of Washington, 105, (2) 366–372.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 1992. "Ephydridae Zetterstedt, 1837 (Insecta, Diptera): Proposed Precedence over Gymnomyzidae Latreille, 1829." The Bulletin of Zoological Nomenclature, 49, (2) 133–136.
  • Mathis, Wayne Neilsen and Edmiston, J. F. 1991. "S. W. Williston's Species of Ephydridae (Diptera) from St. Vincent (West Indies)." Proceedings of the Biological Society of Washington, 104, (4) 816–839.
  • Mathis, Wayne Neilsen and Freidberg, Amnon. 1991. "Review of the Afrotropical Beach Flies of the Tribe Canacini and Subfamily Nocticanacinae (Diptera: Canacidae)." Proceedings of the Entomological Society of Washington, 93, (1) 70–85.
  • Freidberg, Amnon and Mathis, Wayne Neilsen. 1990. "A New Species of Craspedoxantha and a Revised Phylogeny for the Genus (Diptera: Tephritidae)." Proceedings of the Entomological Society of Washington, 92, (2) 325–332.
  • Mathis, Wayne Neilsen. 1990. "A Revision of the Shore-Fly Genus Diphuia Cresson (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 92, (4) 746–756.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 1990. "A Revision of the Western Paleartic Species of Athryoglossa (Diptera: Ephydridae)." Transactions of the American Entomological Society, 116, (1) 103–133.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 1990. "Taxonomic Notes on Ephydridae (Diptera)." Proceedings of the Biological Society of Washington, 103, (4) 891–906.
  • Mathis, Wayne Neilsen. 1989. "Diptera (Insecta) or True Flies of the Pitcairn Group (Ducie, Henderson, Oeno, and Pitcairn Islands)." Atoll Research Bulletin, 327 1–15. https://doi.org/10.5479/si.00775630.327.1.
  • Mathis, Wayne Neilsen. 1989. "A Review the Beach Flies of the Carribean and Gulf of Mexico (Diptera: Canacidae)." Proceedings of the Biological Society of Washington, 102, (3) 590–608.
  • Mathis, Wayne Neilsen and Jin, Z. Y. 1989. "A Review of the Shore Fly Genus Polytrichophora Cresson from Asia (Diptera: Ephydridae)." Proceedings of the Biological Society of Washington, 102, (2) 434–446.
  • Mathis, Wayne Neilsen. 1988. "First Record of the Shore-Fly Genus Placopsidella KertAcsz from North America (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 90, (3) 334–337.
  • Mathis, Wayne Neilsen. 1988. "The First Record of the Genus Procanace Hendel from North America, with the Description of a New Species (Diptera: Canacidae)." Proceedings of the Entomological Society of Washington, 90, (3) 329–333.
  • Mathis, Wayne Neilsen. 1988. "Beach Flies of the Republic of Seychelles (Diptera: Canacidae)." Bulletin of the Washington Biological Society, 8 22–29.
  • Mathis, Wayne Neilsen and Jin, Z. Y. 1988. "A Review of the Asian Species of the Genus Lamproscatella Hendel (Diptera: Ephydridae)." Proceedings of the Biological Society of Washington, 101, (3) 540–548.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 1988. "Studies on the Shore Fly Tribe Dagini (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 90, (1) 106–121.
  • Mathis, Wayne Neilsen and Deeming, J. C. 1987. "A Revision of the Shore Fly Genus Elephantinosoma Becker (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 89, (3) 562–570.
  • Mathis, Wayne Neilsen. 1986. "A Revision of the Subgenus Eremotrichoma Soika of the Shore Fly Genus Allotrichoma Becker (Diptera: Ephydridae)." Isreal Journal of Entomology, 19 127–139.
  • Mathis, Wayne Neilsen and Hogue, C. L. 1986. "Description of a New Species of the Shore Fly Genus Diedrops (Diptera: Ephydridae) from Colombia." Contributions in Science (Los Angeles County Museum), 377 21–26.
  • Mathis, Wayne Neilsen and Steiner, Warren E. 1986. "An Adventive Species of Brachydeutera Loew in North America (Diptera: Ephydridae)." Journal of the New York Entomological Society, 94, (1) 56–61.
  • Freidberg, Amnon and Mathis, Wayne Neilsen. 1985. "On the Feeding Habits of Rhynchopsilopa (Diptera: Ephydridae)." Entomophaga, 30, (1) 13–21.
  • Mathis, Wayne Neilsen. 1985. "A New Psilopine Genus and Species from Isreal with a Recharacterization of the Tribe and Key to Other Psilopine Genera of the Middle East (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 87, (2) 375–380.
  • Mathis, Wayne Neilsen. 1984. "A Revision of the Genus Asmeringa Becker (Diptera: Ephydridae)." Isreal Journal of Entomology, 17 67–79.
  • Mathis, Wayne Neilsen. 1984. "A Revision of the Shore Fly Genus Homalometopus Becker (Diptera: Ephydridae)." Proceedings of the Biological Society of Washington, 97, (2) 253–264.
  • Mathis, Wayne Neilsen. 1984. "Notes on the Shore Fly Genus Diedrops (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 86, (2) 349–353.
  • Mathis, Wayne Neilsen. 1983. "A revision of the genus Dagus Cresson (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 85, (4) 717–726.
  • Mathis, Wayne Neilsen. 1983. "Notes on Brachydeutera Loew (Diptera: Ephydrididae) from North America." Entomological News, 94, (3) 177–180.
  • Mathis, Wayne Neilsen. 1982. "Description of a New Species of Nocticanace Malloch (Diptera: Canacidae) from Sri Lanka, with Notes on Two Related Species." Proceedings of the Entomological Society of Washington, 84, (3) 421–425.
  • Mathis, Wayne Neilsen. 1982. "Canacidae of Israel, with a Revison of the Paleartic Species of the Genus Canace Haliday (Diptera)." Insect Systematics & Evolution, 13 57–66. https://doi.org/10.1163/187631282X00543.
  • Mathis, Wayne Neilsen and Freidberg, Amnon. 1982. "New Beach Flies of the Genus Xanthocanace Hendel with a Review of the Species from the Western Paleartic (Diptera: Canacidae)." Memoirs of the Entomological Society of Washington, 10 97–104.
  • Mathis, Wayne Neilsen and Thompson, F. Christian. 1982. "Editors' preface and acknowledgments." Memoirs of the Entomological Society of Washington., 4–5.
  • Mathis, Wayne Neilsen. 1981. "Proposed Use of the Plenary Powers to Grant Precedence to the Family-Group Name Ephydridae over Hydrelliidae (Diptera)." The Bulletin of Zoological Nomenclature, 38, (3) 201–204.
  • Mathis, Wayne Neilsen and Wirth, W. W. 1981. "Hydrellia tritici (Diptera: Ephydridae), New to Hawaii." Proceedings of the Hawaiian Entomological Society, 23, (3) 371–373.
  • Thompson, F. Christian and Mathis, Wayne Neilsen. 1981. "Haliday's Generic Names of Diptera First Publised in Curtis' A Guide to...British Insects (1837)." Journal of the Washington Academy of Sciences, 70, (2) 80–89.
  • Mathis, Wayne Neilsen and Freidberg, Amnon. 1980. "A New Species of Nesopsilopa from Florida (Diptera: Ephydridae)." The Florida Entomologist, 63, (3) 279–281.
  • Mathis, Wayne Neilsen and Steyskal, G. S. 1980. "A Revision of the Genus Oedoparena Curran (Diptera: Dryomyzidae: Dryomyzinae)." Proceedings of the Entomological Society of Washington, 82, (3) 349–359.
  • Mathis, Wayne Neilsen and Zack, R. S. 1980. "A New Species of Limnella Malloch (Diptera: Ephydridae) from Mt. Rainier, Washington." Entomological News, 91, (3) 65–67.
  • Deonier, D. L., Mathis, Wayne Neilsen, and Regensburg, J. T. 1979. "Natural History and Life-cycle Stages of Notiphila carinata (Diptera: Ephydridae)." Proceedings of the Biological Society of Washington, 91, (4) 798–814.
  • Mathis, Wayne Neilsen. 1979. "Ecuadorian Parascatella (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 81, (4) 602–610.
  • Mathis, Wayne Neilsen and Wirth, W. W. 1979. "Beach flies of Madagascar (Diptera: Canacidae)." Annals of the Natal Museum, 23, (3) 785–796.
  • Mathis, Wayne Neilsen. 1978. "A Synopsis of Neotropical Eleleides Cresson (Diptera: Ephydridae)." Proceedings of the Entomological Society of Washington, 80, (4) 465–472.
  • Mathis, Wayne Neilsen. 1978. "New Synonymy in Ephydridae (Diptera)." Entomological News, 89, (2-3) 105–106.
  • Mathis, Wayne Neilsen. 1978. "A Revision of the Neartic Species of Limnellia Malloch (Diptera: Ephydridae)." Proceedings of the Biological Society of Washington, 91, (1) 250–293.
  • Mathis, Wayne Neilsen and Wirth, W. W. 1978. "Paracanace New Genus near Canaceoides Cresson, Three New Species, and Notes on their Classification (Diptera: Canacidae)." Proceedings of the Entomological Society of Washington, 80, (4) 524–537.
  • Mathis, Wayne Neilsen. 1977. "A Revision of the Genus Rhysophora Cresson with a Key to Related Genera (Diptera: Ephydridae)." Proceedings of the Biological Society of Washington, 90, (4) 921–945.
  • Mathis, Wayne Neilsen. 1977. "A New Synonym and Lectotype Designation in Ephydridae (Diptera)." Proceedings of the Entomological Society of Washington, 79, (4) 537.
  • Mathis, Wayne Neilsen. 1977. "Key to the Neotropical Genera of Parydrinae with a Revision of the Genus Eleleides Cresson (Diptera: Ephydridae)." Proceedings of the Biological Society of Washington, 90, (3) 553–565.
  • Mathis, Wayne Neilsen and Wirth, W. W. 1977. "A New Genus of Psilopine flies (Diptera: Ephydridae) with Notes on its Relationships." Proceedings of the Entomological Society of Washington, 79, (1) 63–74.
  • Mathis, Wayne Neilsen and Wirth, W. W. 1977. "New Synonymy in Ephydridae (Diptera)." Proceedings of the Entomological Society of Washington, 79, (4) 520.
  • Mathis, Wayne Neilsen and Wirth, W. W. 1976. "A New Neotropical Shore Fly Genus with Two New Species (Diptera: Ephydridae)." Pan-Pacific Entomologist, 52, (2) 126–132.
  • Mathis, Wayne Neilsen. 1975. "A Systematic Study of Coenia and Paracoenia (Diptera: Ephydridae)." Great Basin Naturalist, 35, (1) 65–85.
  • Mathis, Wayne Neilsen. 1974. "Lectotype Designation for Notiphila varia Jones (Diptera: Ephydridae)." Entomological News, 85 64–65.
  • Mathis, Wayne Neilsen. 1973. "A Review of the Genus Borboropsis (Diptera: Heleomyzidae)." Pan-Pacific Entomologist, 49, (4) 373–377.
Book Chapter
  • Kirk-Spriggs, Ashley H. and Mathis, Wayne N. 2021. "Diastatidae (incl. Campichoetidae) (diastatid flies)." In Brachycera: Cyclorrhapha, excluding Calyptratae Suricata 8. Kirk-Spriggs, Ashley H. and Sinclair, Bradley J., editors. 2283–2288. Pretoria, South Africa: South African Biodiversity Institute. In Manual of Afrotropical Diptera, 3.
  • Mathis, Wayne N., Zatwarnicki, T., and Irwin, A. 2021. "Ephydridae (shore flies)." In Brachycera: Cyclorrhapha, excluding Calyptratae Suricata 8. Kirk-Spriggs, Ashley H. and Sinclair, Bradley J., editors. 2193–2230. Pretoria, South Africa: South African Biodiversity Institute. In Manual of Afrotropical Diptera, 3.
  • Munari, Lorenzo and Mathis, Wayne N. 2021. "Canacidae (beach flies, surf flies and surge flies)." In Brachycera: Cyclorrhapha, excluding Calyptratae Suricata 8. Kirk-Spriggs, Ashley H. and Sinclair, Bradley J., editors. 2003–2014. Pretoria, South Africa: South African Biodiversity Institute. In Manual of Afrotropical Diptera, 3.
  • Rung, A. and Mathis, Wayne N. 2021. "Aulacigastridae (sap flies)." In Brachycera: Cyclorrhapha, excluding Calyptratae Suricata 8. Kirk-Spriggs, Ashley H. and Sinclair, Bradley J., editors. 1957–1962. Pretoria, South Africa: South African Biodiversity Institute. In Manual of Afrotropical Diptera, 3.
  • Rung, A. and Mathis, Wayne N. 2021. "Periscelididae (dwarf flies)." In Brachycera: Cyclorrhapha, excluding Calyptratae Suricata 8. Kirk-Spriggs, Ashley H. and Sinclair, Bradley J., editors. 1981–1987. Pretoria, South Africa: South African Biodiversity Institute. In Manual of Afrotropical Diptera, 3.
  • Mathis, Wayne Neilsen, Zatwarnicki, Tadeusz, Stuke, Jens-Hermann, and Deeming, John C. 2017. "Order Diptera, family Ephydridae A conspectus on shore-flies from the United Arab Emirates." In Arthropod fauna of the United Arab Emirates. van Harten, Antonius, editor. 636–761. Abu Dhabi: Department of President's Affairs. 6
  • Mathis, Wayne Neilsen. 2010. "Nannodastiidae (nannodastiid beach flies)." In Manual of Central American Diptera. Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, Norman E., and Zumbado, M. A., editors. 1235–1239. Ottawa: NRC Research Press. 2
  • Mathis, Wayne Neilsen. 2010. "Tethinidae (beach flies)." In Manual of Central American Diptera. Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, Norman E., and Zumbado, M. A., editors. 1109–1114. Ottawa: NRC Research Press. 2
  • Mathis, Wayne Neilsen. 2010. "Diastatidae (diastatid flies)." In Manual of Central American Diptera. Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, Norman E., and Zumbado, M. A., editors. 1207–1210. Ottawa: NRC Research Press. 2
  • Mathis, Wayne Neilsen. 2010. "Ephydridae (shore flies)." In Manual of Central American Diptera. Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, Norman E., and Zumbado, M. A., editors. 1211–1233. Ottawa: NRC Research Press. 2
  • Mathis, Wayne Neilsen. 2010. "Coelopidae (kelp flies)." In Manual of Central American Diptera. Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, Norman E., and Zumbado, M. A., editors. 1013–1016. Ottawa: NRC Research Press. 2
  • Mathis, Wayne Neilsen. 2010. "Canacidae (surf flies)." In Manual of Central American Diptera. Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, Norman E., and Zumbado, M. A., editors. 1115–1119. Ottawa: NRC Research Press. 2
  • Mathis, Wayne Neilsen and Rung, Alessandra. 2010. "Periscelididae (periscelid flies)." In Manual of Central American Diptera. Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, Norman E., and Zumbado, M. A., editors. 1087–1092. Ottawa: NRC Research Press. 2
  • Mathis, Wayne Neilsen and Mathis, Dianne. 2008. "Shore flies (Diptera: Ephydridae) of Plummers Island." In The Invertebrate fauna of Plummers Island, Maryland. Contribution XXX to the Natural History of Plummers Island, Maryland.. Brown, John W., editor. 173–191. Washington, D. C.: Bulletin of the Biological Society of Washington.
  • Mathis, Wayne Neilsen and Grimaldi, David A. 2000. "The first Beckeriella Williston from the Afrotropical Region: two new species from Madagascar (Diptera: Ephydridae)." In A floral and faunal inventory of the Parc National de Marojejy, Madagascar: with reference to elevational variation.. Goodman, S. M., editor. 105–115. Chicago: Fieldiana: Zoology 97.
  • Mathis, Wayne Neilsen and Family Periscelididae. 1998. "Family Canacidae." In Contributions to a Manual of Palaearctic Diptera (with special reference to flies of economic importance). Vol. 3. Higher Brachycera.. Papp, László and Darvas, Béla, editors. 251–257. Budapest: Science Herald.
  • Mathis, Wayne Neilsen, Papp, László, and Family Periscelididae. 1998. "Family Periscelididae." In Contributions to a Manual of Palaearctic Diptera (with special reference to flies of economic importance). Vol. 3. Higher Brachycera.. Papp, László and Darvas, Béla, editors. 285–294. Budapest: Science Herald.
  • Mathis, Wayne Neilsen and Zatwarnicki, Tadeusz. 1998. "Family Ephydridae." In Contributions to a Manual of Palaearctic Diptera (with special reference to flies of economic importance). Vol. 3. Higher Brachycera.. Papp, László and Darvas, Béla, editors. 537–570. Budapest: Science Herald.
  • Papp, László and Mathis, Wayne Neilsen. 1998. "Family Nannodastidae." In Contributions to a Manual of Palaearctic Diptera (with special reference to flies of economic importance). Vol. 3. Higher Brachycera.. Papp, László and Darvas, Béla, editors. 309–314. Budapest: Science Herald.
  • Mathis, Wayne Neilsen. 1991. "Classification of the Shore Flies (Diptera: Ephydridae) Past, Present, and Future." In Proceeding of the II International Congress of Dipterology. 209–228.
  • Mathis, Wayne Neilsen. 1989. "99. Family Ephydridae." In Catalog of the Diptera of the Australasian and Oceanian Regions, special publication 86. 639–649.
  • Mathis, Wayne Neilsen. 1989. "56. Family Pseudopomyzidae." In Catalog of the Diptera of the Australasian and Oceanian Regions, special publication 86. 470.
  • Mathis, Wayne Neilsen. 1989. "103. Family Canacidae." In Catalog of the Diptera of the Australasian and Oceanian Regions, special publication 86. 669–670.
  • Mathis, Wayne Neilsen. 1989. "53. Family Cypselosomatidae." In Catalog of the Diptera of the Australasian and Oceanian Regions, special publication 86. 463.
  • Mathis, Wayne Neilsen. 1989. "84. Family Coelopidae." In Catalog of the Diptera of the Australasian and Oceanian Regions, special publication 86. 563–564.
  • Mathis, Wayne Neilsen and Sasakawa, M. 1989. "102. Family Tethinidae." In Catalog of the Diptera of the Australasian and Oceanian Regions, special publication 86. 86.
  • Wirth, W. W., Mathis, Wayne Neilsen, and Vockeroth, J. R. 1987. "98. Ephydridae." In Manual of the Families and Genera of Neartic Diptera Volume 2. 1027–1047.
  • Mathis, Wayne Neilsen. 1982. "Ephydridae." In Aquatic Biota of Mexico, Central America and the West Indies. 469–472.
  • Mathis, Wayne Neilsen and Thompson, F. Christian. 1982. "Editors' preface and acknowledgments." In The flower flies of the West Indies (Diptera, Syrphidae). 4–5. Washington, DC: The Entomological Society of Washington. 9 in Memoirs of the Entomological Society of Washington..
  • Mathis, Wayne Neilsen. 1981. "Ephydridae." In Aquatic Biota of Tropical South America, Part 1: Arthropoda. 312–316.
  • Mathis, Wayne Neilsen. 1979. "Ephydrinae (Diptera: Ephydridae): A new perspective." In in: First Symposium on the Systematics and Ecology of Ephydridae (Diptera). 47–60.
  • Wirth, W. W. and Mathis, Wayne Neilsen. 1979. "A Review of the Ephydridae (Diptera) Living in Thermal Springs." In in: First Symposium on the Systematics and Ecology of Ephydridae (Diptera). 21–45.
Book
Book Review
  • Thompson, F. Christian and Mathis, Wayne Neilsen. 1981. [Book review] "A Catalogue of Diptera of the Afrotropical Region. British Museum (Natural History)." Bulletin of the Entomological Society of America, 27, (3), 230–231.
  • Mathis, Wayne Neilsen. 1979. [Book review] "Record of My Life-Work in Entomology, 1903-1904, by C. R. Osten Sacken, with an appreciation and introductory preface by K. G. V. Smith." Proceedings of the Entomological Society of Washington, 81, (2), 330–331.