Phylogenetics, molecular evolution, systematics, and historical biogeography of Hymenoptera, especially bees and ants;
Coevolutionary patterns and dynamics between Hymenoptera and their symbionts (e.g., other arthropods, fungi, bacteria);
Evolution of sociality in insects;
Phylogenetic theory and methodology, including applications to large-scale evolutionary questions
My research concentrates on increasing our understanding of the origins, diversification, and molecular evolution of social insects, especially ants and bees. I am particularly interested in applying comparative approaches that incorporate evidence from systematics, ecology, genetics, and genomics in order to address specific questions in ant and bee evolution.