B.S., Biology, State University of New York at Stony Brook, 1975.
M.A., Ecology and Evolution, Johns Hopkins University, 1980.
Ph.D., Ecology and Evolution, Johns Hopkins University, 1983.
Research Interests
Systematics, morphology, and evolution of brackish-coastal and freshwater gastropods, particularly the superfamily Rissooidea; aquatic biogeography, focusing on North America; conservation of freshwater mollusks.